Subfamily Orthotylinae

Orthotylinae Van Duzee 1916

4 tribes and ~50 genera in NA(1), 6 tribes with almost 300 genera worldwide(2)(3)

The Orthotylinae have traditionally been diagnosed by their possession of lamellate, apically divergent parempodia. This type of parempodial structure also occurs in the tribe Pilophorini which is placed in the Phylinae on the basis of the structure of the male genitalia. The male and female genitalia of the Orthotylinae have some structural attributes that offer additional evidence suggesting that the group may be monophyletic. These features include vesical spicules in the male and elaborations of the posterior wall in the female.(4)

Worldwide, especially diverse in S. America and Australia(4)