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Subfamily Phylinae

Plant Bug - Plagiognathus delicatus Small bug - Plagiognathus Mirid on Platanus Plant Bug - Plagiognathus Colorful Mirid at lights - Reuteroscopus ornatus Rhinacloa forticornis - female Rhinacloa forticornis? Harmostes? - Amblytylus nasutus
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Heteroptera (True Bugs)
Infraorder Cimicomorpha
Superfamily Miroidea
Family Miridae (Plant Bugs)
Subfamily Phylinae
Synonyms and other taxonomic changes
Author: Dougla and Scott, 1865
Size
4 tribes and 290 genera in the world. (1)
Identification
The Phylinae are diagnosed primarily on the structure of male genitalia, which are distinctive in their possession of a sclerotized, rigid, non-inflatable vesica and the phallotheca attached to the posterior wall of the genital capsule rather than to the phallobase as in all other Miridae. Parempodial structure in the group varies from setiform, to weakly fleshy, to fleshy, recurved, and apically divergent. The posterior wall in the female genitalia is simple and unornamented. (1)
Remarks
* The classification is based on Schuh et al., 2013(2).

Subfamily PHYLINAE
Genera Cyrtopeltocoris, Phoradendrepulus


Tribe Decomini
Genera Aurantiocoris

Tribe Nasocorini
Genera Adenostomocoris, Arctostaphylocoris, Atomoscelis, Atractotomus, Beckocoris, Campylomma, Chlamydatus, Chlamyopsallus, Hambletoniola, Knigntomiroides, Larinocerus, Lattinophylus, Megalopsallus, Monosynamma, Nevadocoris, Phoenicocoris, Pinomiris, Pruneocoris, Rhinacloa, Spanagonicus, Tannerocoris


Tribe Exaeretini
Genera Gonoporomiris, Hyalopsallus, Tuponia

Subtribe Cremnorrhinina
Genera Amblytylus, Brachyceratocoris, Calidroides, Coquillettia, Guentherocoris, Leutiola, Lopus, Macrotylus, Orectoderus, Pronotocrepis, Strophopoda, Teleorhinus, Ticua


Tribe Phylini
Subtribe Keltoniina


Subtribe Phylina
Genera Conostethus, Icodema, Izyaius, Lepidargyrus, Phylus, Psallus, Sthenarus


Subtribe Oncotylina
Genera Americodema, Arizonapsallus, Asciodema, Bisulcopsallus, Cariniocoris, Ceratopsallus, Ceratopsallus, Compsidolon, Coniferocoris(3), Crassomiris, Europiella, Galbinocoris, Hamatophylus, Hoplomachus, Insulaphylus, Knightopiella, Knightopsallus, Lineatopsallus, Maculamiris, Megalocoleus, Occidentodema, Oligotylus, Oncotylus, Parapsallus, Phallospinophylus, Phyllopidea, Phymatopsallus, Piceophylus, Pinophylus, Plagiognathus, Plesiodema, Psallovius, Pygovepres, Quercophylus, Quernocoris, Ranzovius, Rhinocapsus, Rubellomiris, Rubeospineus, Salicopsallus, Sthenaropsidea, Stictopsallus, Vanduzeephylus, Vesperocoris


Tribe Semiini
Subtribe Semiina
Genera Criocoris, Hoplomachidea, Nicholia, Semium, Tytthus


Genera Alepidiella, Pilophorus, Sthenaridea


Subtribe Tuxedoina
Genera Tuxedo




* Adenostomocoris, Arctostaphylocoris, Aurantiocoris, Guentherocoris, Pruneocoris, and Vanduzeephylus were described by Schuh and Schwartz, 2004(4).
* Knightopiella was described by Schuh, 2004(5).
* Calidroides and Chlamyopsallus were described by Schwartz, 2005(6).
* Izyaius was described in Schwartz, 2006(3).
* Galbinocoris, Hamatophylus, Insulaphylus, Maculamiris, Quernocoris, and Vesperocoris were described by C. Weirauch, 2006(7).
* Crassomiris, Phallospinophylus, Quercophylus, Pygovepres, Rubellomiris, and Rubeospineus were described by C. Weirauch, 2006(8).
* Arizonapsallus, Bisulcopsallus, Ceratopsallus, Cercocarpopsallus, Knightopsallus, Salicopsallus, and Stictopsallus were described by Schuh, 2006(9)
* Lattinophylus was described in Schuh, 2008(10).
* Tytthus was moved out of the Leucophoropterini and into a broadly conceived Phylini by Menard & Schuh, 2011(11).
* Coquillettia was transferred from Hallodapini to Phylini by Denise Wyniger, 2011(12).
* Leutiola and Ticua was described by Denise Wyniger, 2011(12).
Internet References
Systematic Catalog with searchable database and navigable directory (Plant Bug Planetary Biodiversity Inventory, American Museum of Natural History)
Works Cited
1.Plant Bug Planetary Biodiversity Inventory
2.A revised classification of the Phylinae (Insecta, Heteroptera, Miridae) : arguments for the placement of genera
Randall T. Schuh and Katrina L Menard. 2013. American Museum of Natural History.
3.Review of Plesiodema Reuter and a description of a new genus to accommodate Psallus sericeus Heidemann ...
Michael D. Schwartz. 2006. KMK Scientific Press.
4.New genera, new species, new synonyms, and new combinations in North America and Caribbean Phylinae (Heteroptera, Miridae)
Randall T. Schuh and Michael D. Schwartz. 2004. American Museum of Natural History.
5.Revision of Europiella Reuter in North America, with the Description of a New Genus (Heteroptera: Miridae: Phylinae)
Randall T. Schuh. 2004. American Museum of Natural History.
6.Redescription of Strophopoda aprica Van Duzee and the Description of Two New Genera and Five New Species from ...
Michael D. Schwartz. 2005. American Museum of Natural History.
7.New genera, new species, and new combinations in western Nearctic Phylini (Heteroptera, Miridae, Phylinae)
Christiane Weirauch. 2006. American Museum of Natural History.
8.New Genera and Species of Oak-Associated Phylini (Heteroptera: Miridae: Phylinae) from Western North America
Christiane Weirauch. 2006. American Museum of Natural History.
9.Revision, phylogenetic, biogeographic, and host analysis of the endemic western North American Phymatopsallus group, ...
Randall T. Schuh. 2006. American Museum of Natural History.
10.Beckocoris Knight, Nevadocoris Knight, and Lattincoris artemisii, new genus and new species, Asteraceae feeding Phylinae
Randall T. Schuh. 2008. Entomological Society of Washington.
11.Revision of Leucophoropterini: ...
Katrina L. Menard & Randall T. Schuh. 2011. American Museum of Natural History.
12.Revision of the Nearctic Genus Coquillettia Uhler With a Transfer to the Tribe Phylini, the Description of 14 New Species, ...
Denise Wyniger. 2011. New York Entomological Society.