Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
Moths Butterflies Flies Caterpillars Flies Dragonflies Flies Mantids Cockroaches Bees and Wasps Walkingsticks Earwigs Ants Termites Hoppers and Kin Hoppers and Kin Beetles True Bugs Fleas Grasshoppers and Kin Ticks Spiders Scorpions Centipedes Millipedes

Upcoming Events

Photos of insects and people from the 2022 BugGuide gathering in New Mexico, July 20-24

National Moth Week was July 23-31, 2022! See moth submissions.

Photos of insects and people from the Spring 2021 gathering in Louisiana, April 28-May 2

Photos of insects and people from the 2019 gathering in Louisiana, July 25-27

Photos of insects and people from the 2018 gathering in Virginia, July 27-29

Photos of insects and people from the 2015 gathering in Wisconsin, July 10-12

Previous events


Subfamily Dytiscinae

1 2 3 4 5 6 7 8 9 ...
next page
last page

By Barman, EH, White, BP, Jackson, R and Wolfe, GW
Florida Entomologist 91(2), pp. 315-316, 2008, 2008
Scientific notes published by Department of Biological & Environmental Sciences, Georgia College & State University, Milledgeville, Georgia, 2008. Identification of larva based on morphology and breeding seasons. Available on internet form Florida Center for Library Automation (FCLA)

Resolving the taxonomic conundrum in Graphoderus of the east Palearctic with a key to all species (Coleoptera, Dytiscidae)
By Sandra Holmgren, Robert B Angus, Fenglong Jia, Zhen-ning Chen, Johannes Bergsten
ZooKeys 574: 113-142, 2016
Full text (PDF]
Key for all species (Neartic and Paleartic)

Taxonomic revision of the Holarctic diving beetle genus Acilius Leach (Coleoptera: Dytiscidae)
By Bergsten, Johannes & Miller, Kelly B.
Systematic Entomology, 31: 145-197, 2006
Link to abstract.
Describes a new species, Acilius confusus for North America and includes a revised key for all species in the genus.

Two New Species of Thermonectus (Coleoptera: Dytiscidae) from Southwestern United States and Western Mexico
By Kenneth L. Goodhue-McWilliams
The Coleopterists Bulletin, Vol.35 No.4 pp.399-407, 1981

Revision of the genus Eretes Laporte, 1833 (Coleoptera: Dytiscidae)
By Miller, Kelly B.
Aquatic Insects 24(4):247-272., 2002
PDF of this and many other aquatic beetle papers available at

Phylogeny and classification of the tribe Hydaticini (Coleoptera: Dytiscidae)...
By Miller K.B., Bergsten J., Whiting M.F.
Zoologica Scripta, 1-25, 2009
Full title: Phylogeny and classification of the tribe Hydaticini (Coleoptera: Dytiscidae): partition choice for Bayesian analysis with multiple nuclear and mitochondrial protein-coding genes
Full text

New Dytiscidae
By Fall, H. C.
Journal of the New York Entomological Society 25(3):163-182., 1917
Description of several dytiscid species, including Hydroporus arizonicus (now in Neoporus).

A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga)
By Nilsson and Hajek
Distributed by authors, 2019

1 2 3 4 5 6 7 8 9 ...
next page
last page