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Subfamily Dytiscinae

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By Barman, EH, White, BP, Jackson, R and Wolfe, GW
Florida Entomologist 91(2), pp. 315-316, 2008, 2008
Scientific notes published by Department of Biological & Environmental Sciences, Georgia College & State University, Milledgeville, Georgia, 2008. Identification of larva based on morphology and breeding seasons. Available on internet form Florida Center for Library Automation (FCLA)

Resolving the taxonomic conundrum in Graphoderus of the east Palearctic with a key to all species (Coleoptera, Dytiscidae)
By Sandra Holmgren, Robert B Angus, Fenglong Jia, Zhen-ning Chen, Johannes Bergsten
ZooKeys 574: 113-142, 2016
Full text (PDF]
Key for all species (Neartic and Paleartic)

Taxonomic revision of the Holarctic diving beetle genus Acilius Leach (Coleoptera: Dytiscidae)
By Bergsten, Johannes & Miller, Kelly B.
Systematic Entomology, 31: 145-197, 2006
Link to abstract.
Describes a new species, Acilius confusus for North America and includes a revised key for all species in the genus.

Two New Species of Thermonectus (Coleoptera: Dytiscidae) from Southwestern United States and Western Mexico
By Kenneth L. Goodhue-McWilliams
The Coleopterists Bulletin, Vol.35 No.4 pp.399-407, 1981

Revision of the genus Eretes Laporte, 1833 (Coleoptera: Dytiscidae)
By Miller, Kelly B.
Aquatic Insects 24(4):247-272., 2002
PDF of this and many other aquatic beetle papers available at

Phylogeny and classification of the tribe Hydaticini (Coleoptera: Dytiscidae)...
By Miller K.B., Bergsten J., Whiting M.F.
Zoologica Scripta, 1-25, 2009
Full title: Phylogeny and classification of the tribe Hydaticini (Coleoptera: Dytiscidae): partition choice for Bayesian analysis with multiple nuclear and mitochondrial protein-coding genes
Full text

A systematic revision of species of Dytiscus Linnaeus (Coleoptera: Dytiscidae). Part 1. Classification based on adult stage
By Roughley R.E.
Quaestiones Entomologicae 26: 383-557, 1990

New Dytiscidae
By Fall, H. C.
Journal of the New York Entomological Society 25(3):163-182., 1917
Description of several dytiscid species, including Hydroporus arizonicus (now in Neoporus).

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