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Suborder Cyphophthalmi - Mite harvestmen

harvestman #6 voucher image - Siro boyerae harvestman #6 voucher image - Siro boyerae Metasiro sassafrasensis Siro exilis Siro exilis Mite harvestman - Siro exilis Mite harvestman - Siro Siro - female
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Chelicerata (Chelicerates)
Class Arachnida (Arachnids)
Order Opiliones (Harvestmen)
Suborder Cyphophthalmi (Mite harvestmen)
Numbers
11 spp. in 2 families in US & Canada (1):
Family Neogoveidae
Metasiro americanus, M. sassafrasensis, M. savannahensis

Family Sironidae
Siro acaroides, S. boyerae, S. calaveras, S. clousi, S. exilis, S. kamiakensis, S. shasta, S. sonoma

~200 spp. in 6 families worldwide(2)
Identification
Mite harvestmen are not likely to be confused with other US + Canada Opiliones, but because of their small size and appearance, can be mistaken for some mites. Look for the distinctive ozophores (anterolateral knob-like structures on cephalothorax, with openings to defensive glands).
Range
western US - Siro
Appalachians (Maryland, West Virginia, Virginia) - Siro exilis
southeastern US - Metasiro
Habitat
Rarely encountered & collected unless specifically targeted. A productive means of collection is to sift leaf litter, using either a mesh sifter or litter concentrator. Forested upland habitats are promising places to search.
Print References
Shear WA. 1980. A review of the Cyphophthalmi of the United States and Mexico, with a proposed reclassification of the suborder (Arachnida, Opiliones). American Museum Novitates, 2705, 1–34. (online PDF)

Giribet G & WA Shear. 2010. The genus Siro Latreille, 1796 (Opiliones, Cyphophthalmi, Sironidae), in North America with a phylogenetic analysis based on molecular data and the description of four new species. Bulletin of the Museum of Comparative Zoology, 160(1), 1-33. (online PDF)

Clouse RM & WC Wheeler. 2014. Descriptions of two new, cryptic species of Metasiro (Arachnida: Opiliones: Cyphophthalmi: Neogoveidae) from South Carolina, USA, including a discussion of mitochondrial mutation rates. Zootaxa, 3814(2), 177­–201. (online PDF)
Internet References
Works Cited
1.Cyphophthalmi Checklist
Giribet, Gonzalo. 2000. Revista Ibérica de Aracnología.
2.Kury A.B. (-2014) Classification of Opiliones