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Photo#256984
Winter Cranefly? - Cramptonomyia spenceri - female

Winter Cranefly? - Cramptonomyia spenceri - Female
Puyallup, Pierce County, Washington, USA
March 6, 2009
Size: around 15 mm

Images of this individual: tag all
Winter Cranefly? - Cramptonomyia spenceri - female Fly - Cramptonomyia spenceri - female Fly - Cramptonomyia spenceri - female

sex
Did you realize you have a female, Lynette? See comments on my fly from Chen.

 
Female
thanks Kevin, I didn't know that.

great job!
another family off the fly wish list.

Size
I have some specimens of this species in the lab and they definitely seem bigger than 5-6 mm. I will measure them tomorrow to make sure, but Lynette's measurements are probably correct.

 
Size II
I looked at the specimens in the lab, and they are definitely bigger than 5-6 mm. They appear to be about 12-13 mm closer to Lynette's 15mm.

 
size
I found one of these the week after Lynette found hers (thanks for stealing my thunder, Lynette!). Mine was also quite large, like hers - definitely NOT 5-6 mm.

 
No problem
:)

Moved
Moved from "Nematocera" (Non-Brachycera).

I have to overlook the size difference because everything else matches as far as I can tell. Ideally somebody will find a collection specimen to compare, or the original description: Alexander, 1931, Bull. Brooklyn Ent. Soc. xxvi, pp. 7-11.

 
Awesome.
I'm glad to add a new family of flies to the guide. I'll be on the lookout for more of these now that I know they aren't winter craneflies.

Arnett
Following the key in Arnett I end up in Anisopdinae but this isn't a very good match for the genera we have in the guide and there aren't any others in North America. (And it's a bit small.)

If I count right, R has three branches, M has three branches, Cu has 2 branches, cell d is present, C ends at R5.

Crampiomyia might fit but is less than half the size of this.

Ocelli rule out several families.

 
I would agree
that it is probably Cramptonomyia spenceri, in the family Pachyneuridae (or Cramptonomyiidae if you're a splitter). The wing venation (especially the crossvein between R2+3 and R4+5 which seems diagnostic) and other features seem to suggest that species (the only one in the genus). Great new family for Bugguide!

 
Four antennae;
are those antennae or really long mouthparts?

 
Palps
I think they are just long palps.

Proboscis
Note that shadow of a long proboscis in addition to antennae. Like a mosquito, but the rest doesn't quite match.

Is the 15mm size body length only? Do you have a closer view of wing veins or a different angle? There's a rare, much smaller gnat not represented in the guide which has a stigma and generally similar veins.

 
15 mm
is tip of head to end of wings. I'll check for other pics.

 
How determined?
How did you come up with 15mm?

The wing venation of Cramptonomyia is a better fit than Anisopodinae but the size is way off. Arnett says 5-6mm.

Was the environment rainforest with alder trees present?

 
Size
was from comparing my fingers to bug, it was the same as the end of my thumb. I can see it being off some, but not by 10 mm.

It certainly is rainy here, but we are not considered a rainforest. We do have alder. It was found in an open field area, that had been recently cleared within the last couple of years.

Hi Lynette
Just notice here is image of an western winter crane fly. Notice the differences of the venation. - Chen

 
Thanks
I was way off!

Moved
Moved from Crane Flies.

Based on the
image there is only one visible anal vein, which will exculde it from both families of crane fly (Tipulidae) and winter crane fly (Trichoceridae). The other families that I can think about that it might fit in are Anisopodidae or Mycetophilidae. Let's wait for other opionions. - Chen

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