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Family Enicocephalidae - Gnat Bugs

tiny insect - Systelloderes biceps - male help with i.d. please - Systelloderes Systelloderes sp., Enicocephalidae - Unique-headed Bug - Systelloderes Bug - Systelloderes Bug - Systelloderes Enicocephalid - Systelloderes biceps strange bug - Systelloderes Immature Nabidae? - Systelloderes
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Heteroptera (True Bugs)
Infraorder Enicocephalomorpha (Unique-headed Bugs)
Family Enicocephalidae (Gnat Bugs)
Other Common Names
Unique-headed Bugs(1)(2)
Numbers
poorly understood group(3) with ~10 spp. in 2 genera in our area and >400 spp. in 55 genera worldwide(4)
in our area: Boreostolus (2 spp., nw. US & ne. Asia): B. americanus Wygodzinsky & Stys 1970 (nw. US); Systelloderes (genus needs revision)
Size
2-5 mm [worldwide, 1-17 mm(5)]
Identification
slender body with unusual elongated head constricted behind eyes and at base; ocelli present; antennae and beak 4-segmented; forewings entirely membranous; front femora and tarsi thickened; middle and hind tarsi 2-segmented.
Range
throughout the world and much of the US
Habitat
under leaf litter, rocks, bark
Food
Predacious(2)
Life Cycle
Some species form swarms in sunny patches in woodland, resembling the behavior of midges(1)
Remarks
may form aggregations of many thousands of individuals(4), but mostly rarely encountered(6)(7)
Works Cited
1.How to Know the True Bugs
Slater, James A., and Baranowski, Richard M. 1978. Wm. C. Brown Company.
2.American Insects: A Handbook of the Insects of America North of Mexico
Ross H. Arnett. 2000. CRC Press.
3.Revision of the New World Enicocephalomorpha (Heteroptera)
Wygodzinsky, P.W. and K. Schmidt. 1991. Bulletin of the American Museum of Natural History ; no. 200.
4.Biodiversity of the Heteroptera
Henry T.J. 2009. In: Foottit R.G., Adler P.H., eds. Insect biodiversity: Science and society. Oxford: Wiley-Blackwell: 223-263.
5.Australian Faunal Directory
6.A Field Guide to Insects
Richard E. White, Donald J. Borror, Roger Tory Peterson. 1998. Houghton Mifflin Co.
7.Borror and DeLong's Introduction to the Study of Insects
Norman F. Johnson, Charles A. Triplehorn. 2004. Brooks Cole.