Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Family Enicocephalidae - Gnat Bugs

tiny insect - Systelloderes biceps - male Systelloderes sp., Enicocephalidae - Unique-headed Bug - Systelloderes Unique-headed Bug - Systelloderes Gnat bug from north central Texas 8851 - Hymenocoris 8851 - Hymenocoris Systelloderes sp. - Systelloderes Systelloderes biceps? - Systelloderes
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Heteroptera (True Bugs)
Infraorder Enicocephalomorpha (Unique-headed Bugs)
Family Enicocephalidae (Gnat Bugs)
Other Common Names
Unique-headed Bugs(1)(2)
Explanation of Names
Enicocephalidae Stål 1860
Numbers
poorly understood group(3) with ~10 spp. in 6 genera in our area and >400 spp. in 55 genera worldwide(4)
Size
1‒17 mm(5); our spp., 2‒5 mm
Identification
slender body with unusual elongated head constricted behind eyes and at base; ocelli present; antennae and beak 4-segmented; forewings entirely membranous; front femora and tarsi thickened; middle and hind tarsi 2-segmented.
Range
worldwide
Habitat
under leaf litter, rocks, bark
Life Cycle
Some species form swarms in sunny patches in woodland, resembling the behavior of midges(1)
Remarks
may form aggregations of many thousands of individuals(4)
Works Cited
1.How to Know the True Bugs
Slater, James A., and Baranowski, Richard M. 1978. Wm. C. Brown Company.
2.American Insects: A Handbook of the Insects of America North of Mexico
Ross H. Arnett. 2000. CRC Press.
3.Revision of the New World Enicocephalomorpha (Heteroptera)
Wygodzinsky, P.W. and K. Schmidt. 1991. Bulletin of the American Museum of Natural History ; no. 200.
4.Biodiversity of the Heteroptera
Henry T.J. 2009. In: Foottit R.G., Adler P.H., eds. Insect biodiversity: Science and society. Oxford: Wiley-Blackwell: 223−263.
5.Australian Faunal Directory