Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

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National Moth Week was July 19-27, and the Summer 2025 gathering in Louisiana, July 19-27

Photos of insects and people from the 2024 BugGuide gathering in Idaho July 24-27

Moth submissions from National Moth Week 2024

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Family Dryinidae - Pincer Wasps

Representative Images

wasp - Deinodryinus atriventris Braconidae Wasp - male Female, Gonatopus? - female dryinid Anteon masoni Dryinidae? Dryinidae

Classification

Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon (Aculeata - Ants, Bees and Stinging Wasps)
Superfamily Chrysidoidea (Cuckoo Wasps & allies)
Family Dryinidae (Pincer Wasps)

Explanation of Names

Dryinidae Haliday, 1833

Numbers

123 spp. in 15 genera and 7 subfamilies in our area;(1)(2) ~2000 spp. in >50 genera and 17 subfamilies worldwide
Subfamily Anteoninae
1. Anteon: 19 spp.
2. Deinodryinus: 4 spp.
3. Lonchodryinus: 3 spp.

Subfamily Aphelopinae
4. Aphelopus: 4 spp.
5. Crovettia: 2 spp.

Subfamily Bocchinae
6. Bocchus: 11 spp.

Subfamily Dryininae
7. Dryinus: 15 spp.

Subfamily Gonatopodinae
8. Echthrodelphax: 2 spp.
9. Esagonatopus: 2 spp.
10. Gonatopus: 52 spp.
11. Neodryinus: 4 spp.
12. Pentagonatopus: 1 sp.
13. Trichogonatopus: 1 sp.

Subfamily Plesiodryininae
14. Plesiodryinus: 1 sp.

Subfamily Thaumatodryininae
15. Thaumatodryinus: 2 spp.

Identification

Females usually have chelate protarsus to hold the host during oviposition.(3) Females of most spp. are wingless; some mimic ants. Antennae 10-segmented. Only the female has front legs with oversized tarsal claws, used to grab leafhoppers.(4)


Male wing venation

Range

worldwide and throughout NA

Food

parasitoids and predators of Auchenorrhyncha (most commonly Cicadellidae, Delphacidae, Flatidae)(3)

Life Cycle

Larva develops inside the host, then extrudes in a characteristic sac bulging out of the host's abdomen. Pupation on host plant or in the soil.(3)
Once the female captures a leafhopper, she inserts her ovipositor into its abdomen. As the larva grows, it moves to a chamber that it forms from its molted skins. This chamber protrudes from the side of the host. This protects it from the immune response of the leafhopper.(4)

Print References

Evenhuis, Neal L. (2005). Bibliography of and New Taxa Proposed by R.C.L. Perkins (1866–1955). Bishop Museum Occasional Papers, 83: 1-55. (Full text)
Guglielmino, Adalgisa; Olmi, Massimo; Marletta, Alessandro; & Speranza, Stefano. (2018). A new species of the genus Gonatopus Ljungh from the USA (Hymenoptera, Dryinidae). ZooKeys, 747: 63–69. (Full text)
Lucchi, Andrea & Wilson, Stephen W. (2003). Notes on Dryinid Parasitoids of Planthoppers (Hymenoptera: Dryinidae; Hemiptera: Flatidae, Issidae). Journal of the Kansas Entomological Society, 76(1): 73-75. (Full text)
Virla, Eduardo G.; Moya-Raygoza, Gustavo; & Guglielmino, Adalgisa. (2023). A review of the biology of the pincer wasps (Hymenoptera: Dryinidae). Austral Entomology, 62(3): 274–299. (Full text)

Internet References

Works Cited

1.Catalog of Hymenoptera in America North of Mexico
Karl V. Krombein, Paul D. Hurd, Jr., David R. Smith, and B. D. Burks. 1979. Smithsonian Institution Press.
2.Nomina Insecta Nearctica
Poole, Robert W. Nearctica.com, Inc.
3.van Noort S. (2004-2015) WaspWeb: Wasps, bees and ants of Africa and Madagascar
4.Hidden Company that Trees Keep: Life from Treetops to Root Tips
James B. Nardi. 2023. Princeton University Press.