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Family Mydidae - Mydas Flies

Delhi Sands Flower-loving Fly - Rhaphiomidas terminatus - male Is this a bee fly? - Mydas atratus fly? - Phyllomydas parvulus - male Three Ring Mydas - Mydas chrysostomus Paratype from the Cal Academy of Sciences - Pseudonomoneura nelsoni - female Mydas clavatus Large Wasp/Fly (?) with Orange Legs/Antennae  - Mydas tibialis Unknown Fly
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Diptera (Flies)
No Taxon (Orthorrhapha)
Superfamily Asiloidea
Family Mydidae (Mydas Flies)
Synonyms and other taxonomic changes
Mydaidae
Explanation of Names
Mydidae Latreille 1809
see Mydas
Numbers
~80 spp. in 9 genera of 4 subfamilies in our area(1), ~500 spp. in 66 genera of 11 subfamilies worldwide(1)(2)
Size
9-60 mm(3) (our spp. to ~30 mm); neotropical Gauromydas heros is world's largest dipteran(1)
Identification
Large flies (often wasp mimics) with prominent, clubbed antennae and distinctive wing venation:

Range
Worldwide, primarily tropical/subtropical, most diverse in Subsaharan Africa (esp. in the south) but poorly represented in the Oriental Region, with some representation in temperate climes(1); in our area, most spp. have restricted distribution, mostly in sw. US(3). Mydas clavatus is widespread, largely eastern.
Habitat
Primarily in arid environments, although quite a few species occur in forests(1), e.g., Mydas clavatus; larvae usually in decaying wood or soil
Season
Summer
Food
larvae of some spp. prey on beetle larvae(3)
Life Cycle
Larva and pupa of Mydas clavatus
Remarks
Batesian mimicry of large spider wasps (Pompilidae: Pepsis, Hemipepsis) by Mydas flies is discussed in(4).
Print References
Artigas, J. N, & N. Papavero (1990). "Studies on Mydidae (Diptera): V. Phylogenetic and biogeographic notes, key to the American genera and illustrations of spermathecae". Gayana Zool. 54(3-4): 87-116 (Full Text)
Hardy, D. E. (1944). New Asilidae and Mydaidae (Diptera) in the Snow Collection. Canad. Ent. 76: 226-230.
Hardy, D.E. (1950). The Nearctic Nomoneura and Nemomydas (Diptera:Mydaidae). Wasmann Journal of Biology 8: 9-37. (Full Text)
Wilcox, J., Papavero, N. (1971). "The American genera of Mydidae (Diptera) with the description of three new genera and two new species(5)". Arquivos Zoologica São Paulo. 21 (2): 41–119 (Full Text)
Wilcox, J., Papavero, N., & T. Pimentel (1989). "Studies of Mydidae (Diptera). IVb. Mydas and allies in the Americas (Mydinae, Mydini)(6)". Museu Paraense "Emilo Goeldi", Coleção Emilie Snethlage, Belém, Brazil. 139 pp.
Works Cited
1.Dikow T. (2001–) Asiloid Flies web-site
2.Order Diptera Linnaeus, 1758. In: Zhang Z.-Q. (ed.) Animal biodiversity: An outline of higher-level classification...
Pape T., Blagoderov V., Mostovski M.B. 2011. Zootaxa 3148: 222–229.
3.American Insects: A Handbook of the Insects of America North of Mexico
Ross H. Arnett. 2000. CRC Press.
4.Sonoran Desert Summer
John Alcock, Marilyn Hoff Stewart. 1994. University of Arizona Press.
5.The American genera of Mydidae (Diptera) with the description of three new genera and two new species
Wilcox, J., Papavero, N. 1971. Arquivos Zoologica (São Paulo, Brazil), Vol. 21 (2):41–119.
6.Studies of Mydidae (Diptera). IVb. Mydas and allies in the Americas (Mydinae, Mydini)
Joseph Wilcox, Nelson Papavero, Therezinha Pimentel. 1989. Museu Paraense "Emilo Goeldi", Coleção Emilie Snethlage, Belém, Brazil.