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Family Pamphiliidae - Webspinning and Leafrolling Sawflies

Cherry Webspinning Sawfly - Neurotoma edwardi Pine sawfly, 12:24pm - Acantholyda erythrocephala - female Sawfly - Onycholyda amplecta Pamphilius ochreipes ? - Pamphilius ochreipes Unknown Sawfly - Pamphilius ochreipes Pamphilius ocreatus? Black and Yellow Sawfly - Onycholyda - male Sawfly
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon ("Symphyta" - Sawflies, Horntails, and Wood Wasps)
Family Pamphiliidae (Webspinning and Leafrolling Sawflies)
Explanation of Names
Pamphiliidae Cameron 1890
Numbers
72 spp. in 5 genera in our area(1)
Identification
Mesonotum not divided by a transverse groove between bases of fore wings. Protibia with 2 apical spurs, the smaller spur at least half as long as larger spur. Fore wing with longitudinal vein Sc between veins C and R (sometimes veins R and Sc touching). Clypeus about as wide as face between eyes; labrum hidden behind clypeus. Abdomen strongly flattened dorsoventrally.
Adults have many-segmented antennae. Larvae with long antennae, long subanal appendages, small hook at apex of last segment. --Dave Smith
keys to spp. in(2)(3)
Food
larvae spin webs in a variety of woody plants where they feed on foliage (see Remarks)
Remarks
The following list (from World Catalog of Symphyta, 2010), with known host plants added (from ECatSym) was provided by Alexander Boldyrev here.

Family PAMPHILIIDAE
Subfamily CEPHALCIINAE
Genus Acantholyda
Subgenus Acantholyda

apicalis (Westwood, 1874)
atripes (Cresson, 1880)
erythrocephala (Linne, 1758) - Pinus
marginiventris (Cresson, 1880)
pini Rohwer, 1911
poeppigii (Zaddach, 1866)
runcinata Middlekauff, 1958
tesselata (Klug, 1808)

Subgenus Itycorsia

aequorea Middlekauff, 1958
albomarginata (Cresson, 1880)
angulata (MacGillivray, 1912)
atrata (Cresson, 1880)
aurigera Middlekauff, 1958
balanata (MacGillivray, 1923)
bicolorata (Norton, 1869)
brunnicans (Norton, 1864)
brunniceps (Cresson, 1880)
bucephala (Cresson, 1880)
burkei Middlekauff, 1958
chicoutimiensis (Huart, 1879)
circumcincta (Klug, 1808)
crocina Middlekauff, 1958
depressa Middlekauff, 1958
floridana Greenbaum, 1975
luteomaculata (Cresson, 1880)
maculiventris (Norton, 1869)
nigripes (Cresson, 1880)
nigrita (Cresson, 1880)
ochrocera (Norton, 1869)
ruficeps (Harrington, 1893)
terminalis (Cresson, 1880)
verticalis (Cresson, 1880)
zappei (Rohwer, 1920)

Acantholyda species not assigned to a subgenus

thalictra Middlekauff, 1958

Genus Cephalcia

californica Middlekauff, 1958
distincta (MacGillivray, 1912)
fascipennis (Cresson, 1880)
frontalis (Westwood, 1874)
fulviceps (Rohwer, 1910)
hopkinsi (Rohwer, 1910)
marginata Middlekauff, 1953
nigra Middlekauff, 1958
provancheri (Huart, 1879)
semidea (Cresson, 1880)

Subfamily PAMPHILIINAE
Genus Neurotoma

crataegi Middlekauff, 1940 - Canada, USA - ?Crataegus
edwardi Liston, 1996 - Canada, USA - ?Padus, ?Prunus
inconspicua (Norton, 1869) - Canada, USA - Prunus
nigra Middlekauff, 1988 - USA - ??
willi Middlekauff, 1958 - USA - ?Crataegus

Genus Onycholyda

amplecta (Fabricius, 1804) - USA - Rubus
excavata (Norton, 1869) - Canada, USA - Cornus
luteicornis (Norton, 1869) - Canada, USA - Rubus
multisignata (Norton, 1864) - Canada, USA - ??
nigritibialis (Rohwer, 1912) - Canada, USA - ??
quebecensis (Provancher, 1878) - Canada, USA - ??
rufofasciata (Norton, 1869) - Canada, USA - ?Dasiphora
sitkensis (Kincaid, 1900) - Canada, USA - ?Rubus

Genus Pamphilius

burquei (Provancher, 1878) - Canada, USA - ??
cinctus Harrington, 1893 - Canada, USA - ??
infuscatus Middlekauff, 1964 - Canada, USA - Populus
middlekauffi Shinohara & D.R. Smith, 1983 - Canada, USA - Cornus, Corylus
ochreipes (Cresson, 1880) - Canada, USA - Viburnum
ocreatus (Say, 1836) - Canada, USA - ??
pacificus (Norton, 1869) - Canada, USA - Rosa
palachei (Ashmead, 1902) - USA - ??
pallimacula (Norton, 1869) - Canada, USA - ?Rosa
persicum MacGillivray, 1907 - Canada, USA - Padus, Persica
phyllisae Middlekauff, 1964 - Canada, USA - ?Quercus
pullatus (Cresson, 1880) - Canada, USA - ?Viburnum
rileyi (Cresson, 1880) - USA (Connecticut to West Virginia, west to Michigan, Iowa, Missouri ) - ?Amelanchier
semicinctus (Norton, 1862) - Canada, USA ( Quebec to North Carolina west to Tennessee and Indiana) - Amelanchier, ?Malus, ?Prunus
uniunguis Middlekauff, 1964 - Canada, USA - ??
vernalis Middlekauff, 1964 - Canada, USA - ??

Unplaced Pamphilius

canadensis Norton, 1869 valid is Lyda canadensis Norton, 1869 - Nearctic region - ??
scripta Say, 1824 valid is Tarpa scripta Say, 1824 - Nearctic region - ??

...
? - non-valuated or questionable
?? - unknown
Works Cited
1.American Insects: A Handbook of the Insects of America North of Mexico
Ross H. Arnett. 2000. CRC Press.
2.The North American sawflies of the genera Acantholyda, Cephalcia, and Neurotoma (Hymenoptera: Pamphiliidae)
Middlekauff W.W. 1958. University of California Publications in Entomology 14: 51-174.
3.The North American sawflies of the genus Pamphilius (Hymenoptera: Pamphiliidae)
Middlekauff W.W. 1964. University of California Publications in Entomology 38: 1-80.