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Family Heteroceridae - Variegated Mud-loving Beetles

Heteroceridae? - Heterocerus fenestratus Tropicus pusillus (Say) - Tropicus pusillus mud beetle #5 - Heterocerus beetle - Heterocerus fenestratus Heterocerus texanus (Pacheco) - Heterocerus texanus Tan-patterned beetle - Heterocerus Pennsylvania Beetle for ID - Heterocerus Tropicus pusillus? - Tropicus pusillus
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga
No Taxon (Series Elateriformia)
Superfamily Byrrhoidea
Family Heteroceridae (Variegated Mud-loving Beetles)
Synonyms and other taxonomic changes
for most recent treatment see (1)
Explanation of Names
Heteroceridae W.S. MacLeay 1825
~40 spp. in 3 genera in our area(2)(1), ~500 spp. in 15 genera worldwide(3)
sandy or muddy shores of rivers, lakes, and ponds
Print References
Pacheco F. (1964) Sistemática, filogenia y distribución de los Heteroceridos de América (Coleoptera: Heteroceridae). Monogràfías del Colegio de Postgraduados, Chapingo, Mexico. 115 pp.
Miller W.V. (1988) A new species of Heterocerus recorded from Florida and Canada (Coleoptera: Heteroceridae). Fla. Entomol. 71: 30‒33. (Full text)
Miller W.V. (1994) New species of Heteroceridae (Coleoptera) from Canada and the United States. Col. Bull. 48: 11‒18. (Full text)
Works Cited
1.Molecular data resolves relationships within Heteroceridae (Coleoptera: Dryopoidea)
King J.G., Starr J.R., Lago P.K. 2011. Syst. Entomol. 36: 435–445.
2.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
3.Order Coleoptera Linnaeus, 1758. In: Zhang Z.-Q. (ed.) Animal biodiversity: An outline of higher-level classification...
Ślipiński S.A., Leschen R.A.B., Lawrence J.F. 2011. Zootaxa 3148: 203–208.
4.The variegated mud-loving beetles (Heteroceridae) of Mississippi and Alabama, with discussion and keys to se US spp.
King, J.G. and P.K. Lago. 2012. Insecta Mundi 0275: 1-53.
5. A catalog of the Coleoptera of America north of Mexico. Family Heteroceridae.
Pacheco, F. 1978. USDA, Agric. Handbook. x + 9 pp. 1978. USDA.
6.Two new species of Heterocerus from North America, with notes on related species (Coleoptera: Heteroceridae).
Miller, W.V. 1988. The Coleopterists Bulletin 42(4): 313-320.