Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
Moths Butterflies Flies Caterpillars Flies Dragonflies Flies Mantids Cockroaches Bees and Wasps Walkingsticks Earwigs Ants Termites Hoppers and Kin Hoppers and Kin Beetles True Bugs Fleas Grasshoppers and Kin Ticks Spiders Scorpions Centipedes Millipedes

Calendar
Upcoming Events

Donate
Giving Tuesday, November 30. Please consider a gift to BugGuide!

BugGuide is a National Moth Week Partner. How to add your National Moth Week 2021 photos. July 17-25.

Photos of insects and people from the Spring 2021 gathering in Louisiana, April 28-May 2

National Moth Week 2020 photos of insects and people.

Photos of insects and people from the 2019 BugGuide Gathering in Louisiana, July 25-27

Discussion, insects and people from the 2018 gathering in Virginia, July 27-29

Photos of insects and people from the 2015 gathering in Wisconsin, July 10-12

Photos of insects and people from the 2014 gathering in Virginia, June 4-7.

Photos of insects and people from the 2013 gathering in Arizona, July 25-28

Previous events


TaxonomyBrowse
Info
ImagesLinksBooksData

Family Heteroceridae - Variegated Mud-loving Beetles

Mud lover (majority) - Heterocerus Hetero 2 - Heterocerus Heteroceridae - Heterocerus pallidus Small unidentified Beetle  - Tropicus pusillus 926C07 - Heterocerus Heterocerus? - Heterocerus 3041642 Heterocerid - Heterocerus 3041764 Heterocerid - Tropicus pusillus
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
No Taxon (Series Elateriformia)
Superfamily Byrrhoidea (Pill, Water and Toe-winged Beetles)
Family Heteroceridae (Variegated Mud-loving Beetles)
Synonyms and other taxonomic changes
for most recent treatment see(1)
Note: The classification proposed in(1) is reflected in the Guide. Summary of changes:
The genus Tropicus is retained in its current circumscription
Species currently treated in Augyles, Centuriatus, Explorator, and Microaugyles all go to the genus Augyles
The remaining New World species go (back!) to Heterocerus
Explanation of Names
Heteroceridae W.S. MacLeay 1825
Numbers
~40 spp. in 3 genera in our area(2)(1), ~500 spp. in 15 genera worldwide(3)
Identification
Range
Widespread
Habitat
sandy or muddy shores of rivers, lakes, and ponds
Print References
(4)(5)(6)
Pacheco F. (1964) Sistemática, filogenia y distribución de los Heteroceridos de América (Coleoptera: Heteroceridae). Monogràfías del Colegio de Postgraduados, Chapingo, Mexico. 115 pp.
Pacheco F. (1969) A new species of Heterocerini (Coleoptera: Heteroceridae). Fla. Entomol. 52: 37-39. (Full text)
Miller W.V. (1988) A new species of Heterocerus recorded from Florida and Canada (Coleoptera: Heteroceridae). Fla. Entomol. 71: 30-33. (Full text)
Miller W.V. (1994) New species of Heteroceridae (Coleoptera) from Canada and the United States. Col. Bull. 48: 11-18. (Full text)
Works Cited
1.Molecular data resolves relationships within Heteroceridae (Coleoptera: Dryopoidea)
J.G. King, J.R. Starr, P.K. Lago. 2011. Systematic Entomology 36(3): 435–445.
2.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
3.Order Coleoptera Linnaeus, 1758. In: Zhang Z.-Q. (ed.) Animal biodiversity: An outline of higher-level classification...
Ślipiński S.A., Leschen R.A.B., Lawrence J.F. 2011. Zootaxa 3148: 203–208.
4.The variegated mud-loving beetles (Heteroceridae) of Mississippi and Alabama, with discussion and keys to se US spp.
King, J.G. and P.K. Lago. 2012. Insecta Mundi 0275: 1-53.
5. A catalog of the Coleoptera of America north of Mexico. Family Heteroceridae.
Pacheco, F. 1978. USDA, Agric. Handbook. x + 9 pp. 1978. USDA.
6.Two new species of Heterocerus from North America, with notes on related species (Coleoptera: Heteroceridae).
Miller, W.V. 1988. The Coleopterists Bulletin 42(4): 313-320.