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Tribe Helopini

 
 
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The Helopini (Coleoptera: Tenebrionidae) of Virginia
By W.E. Steiner, Jr.
Virginia Museum of Natural History Special Publication 16: 331-339, 2009
Full text
The 7 spp. treated represent virtually all the helopine fauna fauna east of the Mississippi River except FL; descriptions & habitus photos are helpful for image-based identification

Doyenellus Nabozhenko and Steiner, a New Genus of Darkling Beetles of the Tribe Helopini from North America
By M.V. Nabozhenko, W.E. Steiner Jr.
Proceedings of the Entomological Society of Washington 123 (3), 564-588, 2021
Full Title: "Doyenellus Nabozhenko and Steiner, a New Genus of Darkling Beetles of the Tribe Helopini (Coleoptera: Tenebrionidae) from North America"

https://doi.org/10.4289/0013-8797.123.3.564

Redefinition of the Opatrine tribes in North America with notes on some apterous genera (Coleoptera:Tenebrionidae:Tenebrionina)
By Rolf L. Aalbu and Charles A. Triplehorn
The Coleopterists Bulletin, Vol. 39, No. 3, pp. 272-280, 1985
Read online free at JSTOR.

A revised subfamily classification of Tenebrionidae (Coleoptera)
By J. C. Watt
New Zealand Journal of Zoology, 1:4, 381-452, 1974

New records of three non-native darkling beetles (Coleoptera: Tenebrionidae) established in California and Nevada, USA
By Steiner W.E., Swearingen J.M.
Col. Bull. 69: 22-26, 2015

Catalogue of Tenebrionidae (Coleoptera) of North America
By Yves Bousquet, Donald B. Thomas, Patrice Bouchard, Aaron D. Smith, Rolf L. Aalbu, M. Andrew Johnston, Warren E. Steiner Jr.
ZooKeys 728: 1-455, 2018
Available online here.

New assignments among the genera Haplandrus LeConte, Metaclisa Jacquelin Du Val and Tharsus LeConte with descriptions...
By Steiner W.E.
Ann. Zool. 66: 529-550, 2016
Full title: New assignments among the genera Haplandrus LeConte, Metaclisa Jacquelin Du Val and Tharsus LeConte with descriptions of larvae and pupae and a new genus for North America (Coleoptera: Tenebrionidae)

https://doi.org/10.3161/00034541ANZ2016.66.4.005

The Tenebrionidae of California: a time sensitive snapshot assessment
By Aalbu R.L., Smith A.D.
ZooKeys 415: 9-22, 2014

 
 
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