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Subfamily Satyrinae - Satyrs, Morphos and Owls

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Notes on the Field Identification of the Intricate Satyr, Hermeuptychia intricata, and Its Ecology in South Carolina
By Thomas Austin
Journal of the Lepidopterists' Society, 72(4):307-314, 2018

Due to strong morphological similarities, the Intricate Satyr Hermeuptychia intricata has been difficult for lepidopterists to visually differentiate from the Carolina Satyr Hermeuptychia sosybius since the former's discovery in 2014. The historical confusion between the two species has resulted in a dearth of information on the ecology and life history of the less abundant and more narrowly distributed H. intricata. I observed adults and larvae of both species in the field at five sites across three counties in the coastal plain of South Carolina, USA.

Subtle Satyrs: differentiation and distribution of the newly described Hermeuptychia intricata in the Southeastern United States
By Andrew Warren, Keith Willmott, and Nick Grishin
News of the Lepidopterists' Society, 56(2) 83-85, 2014
Full text available online.

Summary of publication:
Succinctly displays the morphological distinctions between Hermeuptychia sosybius and H. intricata through both text and figures.

Refining the Diagnostic Characters and Distribution of Hermeuptychia intricata (Nymphalidae: Satyrinae: Satyrini)
By Andrew Warren, Denise Tan, Keith Willmott, and Nick Grishin
Tropical Lepidoptera Research, 24(1): 44-51, 2014
Full text available online.

The absence of androconia on the dorsal surface of the wings is established as an external diagnostic character of male Hermeuptychia intricata Grishin, 2014, that distinguishes this newly described species from males of the sympatric H. sosybius (Fabricius, 1793). Additional United States records of H. intricata are reviewed, extending its distribution to include North Carolina, Georgia, Mississippi, and central Texas. Observations on the phenology and behavior of H. intricata and H. sosybius in northern Florida are given.

A new Hermeuptychia (Lepidoptera, Nymphalidae, Satyrinae) is sympatric and synchronic with H. sosybius ...
By Qian Cong, Nick V. Grishin
ZooKeys, 379: 43–91, 2014
Full title:
A new Hermeuptychia (Lepidoptera, Nymphalidae, Satyrinae) is sympatric and synchronic with H. sosybius in southeast US coastal plains, while another new Hermeuptychia species – not hermes – inhabits south Texas and northeast Mexico.

Abstract & PDF

Confirmation of Rhopalocera (Pieridae, Nymphalidae) previously recorded for Texas and the United States.
By Kendall, R.O.
Journal of the Lepidopterists' Society 28(3): 249-252., 1974
Full Text

Kendall, R.O. 1974. Confirmation of Rhopalocera (Pieridae, Nymphalidae) previously recorded for Texas and the United States. Journal of the Lepidopterists' Society 28(3): 249-252.

The object of this paper is to remove the dubious status of earlier reports of two species of Lepidoptera being found in Texas. Each species is represented at present by a single example only. Examples of earlier recordings have not been found; it is possible, however, that they do exist.

These species may represent single-brooded migrants which come to Texas from time to time.

The Tent Caterpillars
By Terrence D. Fitzgerald
Cornell University Press, 1995
All aspects of the biology of tent caterpillars.

ISBN-13: 978-0801424564
ISBN-10: 0801424569

Butterflies of Oregon, Their Taxonomy, Distribution, and Biology
By Andrew D. Warren
C.P. Gillette Museum of Arthropod Diversity, Dept. of Bioagricultural Sciences and Pest Management, Colorado State University, 2005
408 pages

Book / paper is referenced extensively by other authors (Pyle, James, Nunnallee, etc.)
in their butterfly books about Cascadia / Pacific Northwest.

A Swift Guide to Butterflies of North America: Second Edition
By Jeffrey Glassberg
Princeton University Press, 2017
Publisher's Page
This is a revised second edition of the most detailed, comprehensive, and user-friendly photographic field guide to the butterflies of North America. Written by Jeffrey Glassberg, the pioneering authority on the field identification of butterflies, the guide covers all known species, beautifully illustrating them with 3,500 large, gorgeous color photographs—the very best images available. This second edition includes more than 500 new photos and updated text, maps, and species names. For most species, there are photographs of topsides and undersides, males and females, and variants. All text is embedded in the photographs, allowing swift access in the field, and arrows point to field marks, showing you exactly what to look for. Detailed, same-page range maps include information about the number of broods in each area and where strays have been recorded. Color text boxes highlight information about habitat, caterpillar food plants, abundance and flight period, and other interesting facts. Also included are a quick visual index and a caterpillar food plant index. The result is an ideal field guide that will enable you to identify almost every butterfly you see.

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