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For Insects, Spiders & Their Kin
For the United States & Canada

Species Tomarus sallaei

Scarab beetles of the genus Bothynus in the United States (Coleoptera: Scarabaeidae)
By O. Cartwright
Proceedings of the United States National Museum, 108(3409): 515-541, 1959
Online here.

Monograph of the genus Tomarus (=Bothynus, Ligyrus)

Collagenus dasysternus ... with a key to the New World genera of Pentodontini
By Brett Ratcliffe and Martin Hardy
The Coleopterists Bulletin, Vol. 59, pp.143-150, 2005
Brett Ratcliffe and Martin Hardy, Collagenus dasysternus, a new genus and species of Dynastinae from eastern Venezuela with a key to the New World genera of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 59(1): 143-150, 2005.

From BG's Pentodontini Info tab, "One error noticed; couplet 11- range of genus Aphonus should be 'United States east of the Rocky Mountains'".

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The species of Scarabaeidae (Coleoptera) associated with sugarcane in south Florida
By R.D. Gordon and D.M. Anderson
The Florida Entomologist, 64 (1): 119-138 , 1981
"Those species of Scarabaeidae known to be associated with sugarcane in the Lake Okeechobee area of Florida are discussed and delineated. The species are: Phyllophaga latifrons (LeConte), Anomala marginata (F.), Cyclocephala parallela Casey, Dyscinetus morator (F.), Ligyrus subtropicus (Blatchley), and Euphoria sepulchralis (F.). Descriptions and illustrations of the larvae of all these species are presented as well as descriptions and illustrations of the adults of all species except Euphoria sepulchralis (F.). A key to the south Florida species of the genus Cyclocephala is also included."

A catalog of the Coleoptera of America North of Mexico: Family Scarabaeidae, Subfamilies Rutelinae and Dynastinae
By Hardy A.
USDA Agric. Handbook 529-34b, 57 pp., 1991

The Scarab Beetles of Florida
By Robert Woodruff
Florida Department of Agriculture and Consumer Services, 1973
Full title: Arthropods of Florida and Neighboring Land Areas. Volume 8. The Scarab Beetles of Florida (Coleoptera: Scarabaeidae). Part I. The Laparosticti (Subfamilies: Scarabaeinae, Aphodiinae, Hybosorinae, Ochodaeinae, Geotrupinae, Acanthocerinae). Part II covers May Beetles (Phyllophaga).

Has black-and-white photos. Apparently out of print, but still available from some suppliers--see comments. Also available on-line:

Link updated 7 February 2012--thanks for the comments!

Saprophagous Scarabaeidae (Coleoptera) of North Dakota
By Helgesen R.G., Post R.L.
Schafer-Post Series: North Dakota Insects Publication 7, NDSU. 60 pp., 1967

The phytophagous Scarabaeidae and Troginae (Coleoptera) of North Dakota
By Lago P.K., Post R.L., Oseto C.Y.
Schafer-Post Series: North Dakota Insects Publication 12, NDSU. 131 pp., 1979

The Scarabaeoid beetles of San Diego County, California. Part I. Introduction...
By McPeak R.H., Oberbauer T.A.
Proceedings of the San Diego Society of Natural History 40: 1-24, 2008
Full title: The Scarabaeoid beetles of San Diego County, California. Part I. Introduction and diagnosis of families Glaresidae, Trogidae, Pleocomidae, Geotrupidae, Ochodaeidae, Hybosoridae, and Glaphyridae

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