Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
Moths Butterflies Flies Caterpillars Flies Dragonflies Flies Mantids Cockroaches Bees and Wasps Walkingsticks Earwigs Ants Termites Hoppers and Kin Hoppers and Kin Beetles True Bugs Fleas Grasshoppers and Kin Ticks Spiders Scorpions Centipedes Millipedes

Upcoming Events

BugGuide is a National Moth Week Partner. How to add your National Moth Week 2021 photos. July 17-25.

Photos of insects and people from the Spring 2021 gathering in Louisiana, April 28-May 2

National Moth Week 2020 photos of insects and people.

Photos of insects and people from the 2019 BugGuide Gathering in Louisiana, July 25-27

Discussion, insects and people from the 2018 gathering in Virginia, July 27-29

Photos of insects and people from the 2015 gathering in Wisconsin, July 10-12

Photos of insects and people from the 2014 gathering in Virginia, June 4-7.

Photos of insects and people from the 2013 gathering in Arizona, July 25-28

Previous events


Species Proteoteras willingana - Boxelder Twig Borer - Hodges#3232

3232 Proteoteras willingana  - Proteoteras willingana Green tort moth - Proteoteras willingana 3232 Boxelder Twig Borer  - Proteoteras willingana boxelder twig swelling - Proteoteras willingana A Tortricid Moth - Proteoteras willingana - female Tortricidae, Boxelder Twig Borer, dorsal - Proteoteras willingana Boxelder Twig Borer - Hodges#3232 - Proteoteras willingana Olethreutinae, Boxelder Twig Borer, pupa, dorsal - Proteoteras willingana
Show images of: caterpillars · adults · both
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Lepidoptera (Butterflies and Moths)
Superfamily Tortricoidea (Tortricid Moths)
Family Tortricidae (Tortricid Moths)
Subfamily Olethreutinae
Tribe Eucosmini
Genus Proteoteras
Species willingana (Boxelder Twig Borer - Hodges#3232)
Hodges Number
Synonyms and other taxonomic changes
Proteoteras willingana (Kearfott, 1904)
Proteopteryx willingana Kearfott, 1904
Phylogenetic sequence #621135
Explanation of Names
Specific epithet for Thomas Nathaniel Willing (1858-1920) "in whose honour the species is named." Willing came west as a surveyor and fought in the Battle of Frenchman's Butte. He was appointed Game Warden and Inspector of Weeds for the Northwest Territories. From 1910-1920 he was University of Saskatchewan assistant professor of Natural History where he established the first collections. Kearfott identified and described several species collected by Willing. (1)
Wingspan 18-20 mm (Kearfott, 1904).
Records from Alberta, Saskatchewan and Quebec, Maine south to Tennessee, west to Oklahoma and Iowa. (2), (1)
Larvae feed mostly on Boxelder (Acer negundo), but have been reported on Red Maple (Acer rubrum) . First they skeletonize the undersides of the leaves, then they bore into twigs and buds, which causes a spindle-shaped gall. (3)
Life Cycle
Life cycle images:
Twig gall; larva inside gall; adult
Print References
Kearfott, W.D., 1904. A new Proteopteryx. The Canadian Entomologist 36(10): 306.
Works Cited
1.Assiniboia Micro-Lepidoptera, collected by Mr. T. N. Willing
W. D. Kearfott. 1905. The Canadian Entomologist 37(3): 89-93.
2.North American Moth Photographers Group
3.Maple feeding Tortricidae from the northeastern United States
R. R. Grehan, M. Sabourin, P. M. Hason. 1995. Agricultural Experiment Station, University of Vermont 117: 1-46.