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Subfamily Scymninae

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Survey of lady beetles (Coleoptera: Coccinellidae) found on Texas high plains cotton, alfalfa and weeds.
By Carroll, S.C., M.N. Parajulee, and M.D. Arnold.
Proceedings of the World Cotton Conference (September 10–14, 2007), Lubbock, TX., 2007
Full PDF - Manuscript

A study was conducted during the February to November periods of 2004 and 2005 to determine the seasonal presence and reproduction of lady beetle species on cotton, alfalfa, plus over 20 prevalent Texas High Plains weed species. Over the 2-year study period, a total of 221,209 plant sweeps yielded 27,058 lady beetle adults and 3,936 larvae.

The adult specimens represented a total of six different species with the convergent lady beetle, Hippodamia convergens Guerin-Meneville, easily representing over 80% of the overall population.

Determinate evolution in the color-pattern of the lady-beetles.
By Johnson, R.H.
Carnegie Institution of Washington, Publication No. 122. iv + 104 pp., 1910
Full Text

All of the American coccinellids which could be obtained in numbers and which showed a variable color-pattern were studied, viz, Hippodamini, Coccinellae, and Epilachnini, divisions employed by Casey (1899).

Johnson, R.H. 1910. Determinate evolution in the color-pattern of the lady-beetles. Carnegie Institution of Washington, Publication No. 122. iv + 104 pp.

A checklist of the Coccinellidae of Arkansas.
By Rouse, E.P., and J.B. Chapin.
Proceedings of the Arkansas Academy of Science 30: 76-77., 1976
Full PDF

Rouse, E.P., and J.B. Chapin. 1976. A checklist of the Coccinellidae of Arkansas. Proceedings of the Arkansas Academy of Science 30: 76-77.

The list consists of 49 species and includes distribution by county and ecological information if known.

See also:

Rouse, E.P. and J.B. Chapin. 1980. Checklist of the Lady Beetles (Coccinellidae) of Arkansas. Arkansas Academy of Science, Arkansas Biota Survey Checklist No. 26. 2 pp.

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Adventive lady beetles (Coleoptera: Coccinellidae) in the Canadian Maritime Provinces...
By Hoebeke E.R., Wheeler A.G.
Entomological News 107: 281-290, 1996
Full title: Adventive lady beetles (Coleoptera: Coccinellidae) in the Canadian Maritime Provinces, with new eastern US records of Harmonia quadripunctata

Full text

The Coccinellidae (Coleoptera) of the Maritime Provinces of Canada: new records, biogeographic notes, and conservation concerns
By Majka C.G., McCorquodale D.B.
Zootaxa 1154: 49–68, 2006

Ladybird beetles (Coleoptera: Coccinellidae) recently immigrant to Florida
By Thomas M.C., Blanchard O.J.
Florida Dept. Agric. & Consumer Services Division of Plant Industry, Entomology Circular Number 428. 5 pp., 2013

An annotated checklist of the lady beetles (Coleoptera: Coccinellidae) of Iowa, U.S.A.
By Hesler L.S.
Insecta Mundi 0091: 1-10, 2009

Field guide to recently introduced species of Coccinellidae (Coleoptera) in North America
By Robert Gordon and Natalia Vandenberg
Proceedings of the Entomological Society of Washington, 1991
Full citation:
Gordon, R., and Vandenberg, N. 1991. Field guide to recently introduced species of Coccinellidae (Coleoptera) in North America, with a revised key to North American genera of Coccinellini. Proceedings of the Entomological Society of Washington 93:845-864.

Available online: http://biostor.org/reference/56473

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