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variabilis complex

Scirtidae - Marsh Beetle - Contacyphon Scirtidae - Marsh Beetle - Contacyphon Scirtidae - Marsh Beetle - Contacyphon tan beetle 3 - Contacyphon Fuzzy little beetle - Contacyphon little brown beetle - Contacyphon Cyphon? - Contacyphon Marsh Beetle - Contacyphon
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
No Taxon (Series Elateriformia)
Superfamily Scirtoidea (Plate-thigh and Marsh Beetles)
Family Scirtidae (Marsh Beetles)
No Taxon (former Cyphon)
Genus Contacyphon
No Taxon variabilis complex
Synonyms and other taxonomic changes
Contacyphon variabilis (Thunberg)
Orig. Comb: Cantharis variabilis Thunberg 1785
Syn: Cyphon variabilis (Thunberg)
Syn: Helodes modesta LeConte 1853
Syn: Helodes nebulosa LeConte 1853, but see: Tetrault (1967) (1)
Syn: Helodes punctata LeConte 1853
plus others
Contacyphon variabilis (Thunberg 1787) is holarctic per(2); according to this source, Cyphon modestus (LeConte 1853), Cyphon nebulosus (LeConte 1853), and Cyphon punctatus (LeConte 1853) are all its synonyms.
However, Tetrault (1967) noted distinct genitalic differences and restored the following spp. to full species status: Cyphon nebulosus and Cyphon punctatus. (1)
Print References
Tetrault, R.C. 1967. A revision of the family Helodidae (Coleoptera) for America north of Mexico. Ph.D. thesis, University of Wisconsin. 160 pp.
Works Cited
1.The Water Beetles of Florida
Epler J.H. 2010. Florida Department of Environmental Protection. Tallahassee. 414 pp.
2.Contacyphon Gozis, 1886 removed from synonymy (Coleoptera: Scirtidae)...
Zwick P., Klausnitzer B., Ruta R. 2013. Entomologische Bl├Ątter und Coleoptera 109: 337-353.