Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
Moths Butterflies Flies Caterpillars Flies Dragonflies Flies Mantids Cockroaches Bees and Wasps Walkingsticks Earwigs Ants Termites Hoppers and Kin Hoppers and Kin Beetles True Bugs Fleas Grasshoppers and Kin Ticks Spiders Scorpions Centipedes Millipedes

Calendar
Upcoming Events

Information, insects and people from the 2019 BugGuide Gathering in Louisiana, July 25-27

Discussion, insects and people from the 2018 gathering in Virginia, July 27-29

Photos of insects and people from the 2015 gathering in Wisconsin, July 10-12

Photos of insects and people from the 2014 gathering in Virginia, June 4-7.

Photos of insects and people from the 2013 gathering in Arizona, July 25-28

Photos of insects and people from the 2012 gathering in Alabama

Photos of insects and people from the 2011 gathering in Iowa


TaxonomyBrowse
Info
ImagesLinksBooksData

variabilis complex

Marsh Beetle - Contacyphon Marsh Beetle - Contacyphon tan beetle - Contacyphon Fuzzy little beetle - Contacyphon Beetle - Contacyphon Tiny Beetle - Contacyphon Marsh Beetle - Contacyphon Cyphon - Contacyphon
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
No Taxon (Series Elateriformia)
Superfamily Scirtoidea (Plate-thigh and Marsh Beetles)
Family Scirtidae (Marsh Beetles)
No Taxon (former Cyphon)
Genus Contacyphon
No Taxon variabilis complex
Synonyms and other taxonomic changes
Contacyphon variabilis (Thunberg)
Orig. Comb: Cantharis variabilis Thunberg 1785
Syn: Cyphon variabilis (Thunberg)
Syn: Helodes modesta LeConte 1853
Syn: Helodes nebulosa LeConte 1853, but see: Tetrault (1967) (1)
Syn: Helodes punctata LeConte 1853
plus others
Remarks
Contacyphon variabilis (Thunberg 1787) is holarctic per(2); according to this source, Cyphon modestus (LeConte 1853), Cyphon nebulosus (LeConte 1853), and Cyphon punctatus (LeConte 1853) are all its synonyms.
However, Tetrault (1967) noted distinct genitalic differences and restored the following spp. to full species status: Cyphon nebulosus and Cyphon punctatus. (1)
Print References
Tetrault, R.C. 1967. A revision of the family Helodidae (Coleoptera) for America north of Mexico. Ph.D. thesis, University of Wisconsin. 160 pp.
Works Cited
1.The Water Beetles of Florida
Epler J.H. 2010. Florida Department of Environmental Protection. Tallahassee. 414 pp.
2.Contacyphon Gozis, 1886 removed from synonymy (Coleoptera: Scirtidae)...
Zwick P., Klausnitzer B., Ruta R. 2013. Entomologische Bl├Ątter und Coleoptera 109: 337-353.