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Genus Heterocerus

Variegated Mud-loving Beetle - Heterocerus mollinus Hetero 2 - Heterocerus mud beetle #4 - Heterocerus Hetero 4 - Heterocerus Beetle IMG_9650 - Heterocerus mollinus Heterocerus sp? - Heterocerus gemmatus Heterocerus sp? - Heterocerus gemmatus Heterocerid of sorts - Heterocerus
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
No Taxon (Series Elateriformia)
Superfamily Byrrhoidea (Pill, Water and Toe-winged Beetles)
Family Heteroceridae (Variegated Mud-loving Beetles)
Genus Heterocerus
Synonyms and other taxonomic changes
the genus was split into multiple genera by Pacheco; his classification was adopted in(1)(2)(3), but rejected based on recent analysis(4)(5), with 11 genera erected by Pacheco sunk back to Heterocerus(5)
Explanation of Names
Heterocerus Fabricius 1792
29 spp. in our area, ~200 total(5)
2.6-6.9 mm(5)
all continents except Antarctica(5)
wide variety of riparian habitats; precise habitat does seem to correlate with species composition at a given location and quite different species were often taken from separate habitats that were physically near one another(5)
Print References
Pacheco F. (1964) Sistematica, filogenia, y distribucion de los Heteroceridos de America (Coleoptera: Heteroceridae). Monografias del Colegio de Post-Graduados, Chapingo, Mexico. xii + 155 pp.
Works Cited
1. A catalog of the Coleoptera of America north of Mexico. Family Heteroceridae.
Pacheco, F. 1978. USDA, Agric. Handbook. x + 9 pp. 1978. USDA.
2.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
3.Checklist of beetles (Coleoptera) of Canada and Alaska. Second edition
Bousquet Y., Bouchard P., Davies A.E., Sikes D.S. 2013. ZooKeys 360: 1–402.
4.Molecular data resolves relationships within Heteroceridae (Coleoptera: Dryopoidea)
J.G. King, J.R. Starr, P.K. Lago. 2011. Systematic Entomology 36(3): 435–445.
5.The variegated mud-loving beetles (Heteroceridae) of Mississippi and Alabama, with discussion and keys to se US spp.
King, J.G. and P.K. Lago. 2012. Insecta Mundi 0275: 1-53.