Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Lepidoptera (Butterflies and Moths)
Superfamily Geometroidea (Geometrid and Swallowtail Moths)
Family Geometridae (Geometrid Moths)
Other Common Names
Many species have the word Angle in their common name.
Synonyms and other taxonomic changes
Macaria Curtis, 1826
Eutropa Hübner, 
Philobia Duponchel, 1829
Sciagraphia Hulst, 1896
Pseudoisturgia Povolny & Moucha, 1957
Itame (in part)
Semiothisa (in part)
* In 2008, various species were transferred into Speranza, Letispe, Epelis,
Explanation of Names
Generic epithet Macaria
is from Greek mythology, the daughter of Hercules. (3)
The genus Speranza
is treated as a synonym of Macaria
, Pohl et al. (2016)(5)
and Pohl et al. (2018)(6)
following Sihvonen & Skou (2015:142)(1)
, reversing Ferguson (2008)(2)
More than 100 species in North America, listed under Itame
. 42 species in Canada (CBIF
Wingspan 18-35 mm (except in flightless females that have greatly reduced wings).
Adults of most species have light gray or brown wings (a few are yellow or white) usually with dark markings along the costa, and a dark discal spot in a number of species; some species are unmarked, but many have lines and speckling that ranges from dark and heavy to pale and indistinct - the range of variation in pattern and color makes the genus difficult to describe in general terms.
Represented throughout North America.
Coniferous, mixed, or deciduous forests, plantations, parklands, and woodlots.
Adults usually fly from May to September.
Larvae of many species feed on leaves of coniferous trees; some species feed on broad-leaved trees or shrubs, and a few on herbaceous plants
One generation per year, occasionally two.
Many common species are listed under Itame and Semiothisa in older publications.
Scoble, M.J. (Ed) 1999. Geometrid Moths of the World. A Catalogue.
NHM Publishing, London. 1312 pp.
pinned adult images
of all 42 species in Canada (CBIF)
name change examples
(Dalton State College, Georgia)