Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Subspecies Tropisternus lateralis nimbatus


Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in GSMNP...
By Ferro M.L., Gimmel M.L., Harms K.E., Carlton C.E.
Insecta Mundi 0260: 1–80, 2012
Full title: Comparison of Coleoptera emergent from various decay classes of downed coarse woody debris in Great Smoky Mountains National Park, USA
Full text

Beetle fauna of the United States and Canada
By Marske K.A., Ivie M.A.
The Coleopterists Bulletin 57(4): 495-503, 2003

Illustrated Beetle (Coleoptera) Inventory of the Sabal Palm Sanctuary (Cameron County, Texas)
By Edward G. Riley & Michael A. Quinn
Texas A&M University, Dept. Entomology, College Station, 2011
1,089 species on the SPS list, approximately 14.3% of the documented Texas beetle fauna

Illustrated Beetle (Coleoptera) Inventory of the Sabal Palm Sanctuary (Cameron County, Texas)
Edward G. Riley & Michael A. Quinn
Version Date: January, 2011

Sabal Palm Sanctuary, Brownsville, southmost TX

Classification of the Coleoptera of North America
By LeConte and Horn
Smithsonian Miscellaneous Collections 26(4): i-xxxvii + 1-567., 1883
Full Text

LeConte, J.L. and G.H. Horn. 1883. Classification of the Coleoptera of North America. Prepared for the Smithsonian Institution. Smithsonian Miscellaneous Collections 26(4): i-xxxvii + 1-567.

The aquatic Coleoptera of North Carolina: a biologist's handbook with standard taxonomic effort levels. Version 2.1
By Beaty S.R.
North Carolina Department of Environment and Natural Resources. 56 + 12 pp., 2011

The comparative anatomy of the male genital tube in Coleoptera
By David Sharp
Nabu Press (reproduction), 1912
From the Transactions of the Entomological Society of London, pt. 3 (Dec.) 1912.

Full Text

The water beetles of Maine: including the families Gyrinidae, Haliplidae, Dytiscidae, Noteridae and Hydrophilidae
By Stanley E Malcom
University of Maine Life Sciences and Agricultural Experiment Station Technical Bulletin 48, 1971
PDF version here.

Beetles of Arizona: A Visual Guide with 230 Photos
By Copeland, D.D
Self published, Glendale, California. 60 pp., 2011
Copeland, D.D. 2011. Beetles of Arizona: A Visual Guide with 230 Photos. Self published, Glendale, California. 60 pp.

Most beetles from Madera Canyon and the Santa Rita Mountains south of Tucson.

BioQuip blurb: This convenient guide provides the initial step in identifying beetles found in the Santa Rita Mountains, Huachuca Mountains and other areas in Arizona. Scientific and common names, length, and area or county location are noted for each beetle.

All insects are photographed without any shadows. This allows the focus of attention to be on the characteristics of the beetle and to avoid other background visual distractions. Many beetles have three views, i.e., top, side and front. Most of the photos are from Insects of the Southwest, also produced by Douglas Copeland.