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Photo#385632
Odd Fly - Macrorrhyncha coxalis

Odd Fly - Macrorrhyncha coxalis
Santa Clarita, Los Angeles County, California, USA
April 15, 2010
Size: 3/8 - 1/2 inch

Images of this individual: tag all
Odd Fly - Macrorrhyncha coxalis Odd Fly - Macrorrhyncha coxalis Odd Fly - Macrorrhyncha coxalis

Asindulum?
If this is M. coxalis, it's way out of it's range (cf. http://www.bishopmuseum.org/research/pbs/pdf/BE13-kero-catr.pdf), the northeastern United states. The only two Keroplatidae genera in NA with elongate snouts like that are Macrorrhyncha and Asindulum. The species A. montanum has been found previously in Alta, UT (Cf. Cole 1969), so it is more likely that genus.

Then again, could be a new species of either genus.

In any case, awesome photographs!

 
Johannsen key
I don't see a strong anal vein, which suggests Macrorrhyncha in the key in Manual of Nearctic Diptera. The only species reference I have is Johannsen 1909 which covers all three species of both genera (formerly placed all in Asindulum). Based on his key I would call this coxale.

There are so few records of the species involved that I didn't give heavy weight to recorded range.

 
On second look,
I agree with your assessment of the anal vein, and looked back to Johannsen and it does match the description of M. coxale (M. coxalis in the catalog).

These genera need strong revision in North America before I'll trust the species identification, though.

 
Name changed
I assume coxale vs. coxalis is a mandatory sex change on genus reassignment so I changed the page.

Most of the fungus gnats are in need of revision, I think.

Moved

Moved
Moved from Bibionomorpha. Asindulum?

Maybe Keroplatidae
This may be a member of Keroplatidae. In the picture you added the inner dark vein makes an abrupt turn to the wing margin -- that is a fork (the other branch is not as dark) that is present only in a few kinds of fungus gnat.

See the vein labeled "R4" here: http://commons.wikimedia.org/wiki/File:Keroplatidae_wing_veins.svg

Top view?
Do you have a photo from more directly above, showing the veins in the wings?

 
Added one that's a little bet
Added one that's a little better. I can't really see veins very well.

Moved
Moved from ID Request.

A fungus gnat.

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