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Species Harmonia axyridis - Asian Lady Beetle

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Causes and consequences of ladybug washups in the Finger Lakes region of New York State (Coleoptera: Coccinellidae).
By Denemark, E. and J. Losey.
Entomologica Americana, 116: 78-88., 2010
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Denemark, E. and J. Losey. 2010. Causes and consequences of ladybug washups in the Finger Lakes region of New York State (Coleoptera: Coccinellidae). Entomologica Americana, 116: 78-88.

Abstract.—We searched for and collected data on a phenomenon known as ladybug washups, in which large numbers of coccinellids aggregate on the shores of major bodies of water. Our field season lasted from 5/23/2008 until 8/12/2008 in the Finger Lakes Region of New York, United States.

Aggregation of lady beetles on the shores of lakes (Coleoptera: Coccinellidae).
By Lee, R.E., Jr.
American Midland Naturalist, 104(2): 295-304., 1980
Full PDF

Lee, R.E., Jr. 1980. Aggregation of lady beetles on the shores of lakes (Coleoptera: Coccinellidae). American Midland Naturalist, 104(2): 295-304.

ABSTRACT: Aggregations of lady beetles, predominately Hippodamia convergens and H. tredecimpunctata, were commonly observed on the shores of lakes in the Upper Midwest during the autumn and spring. The beetles remain on the shore for only a short time, usually dispersing within 2-3 weeks. Lady beetles from autumn shore aggregations and overwintering aggregations are characterized by the presence of large amounts of fat, reproductive inactivitiy, empty digestive tracts, a skew in the sex ratio favoring females and the behavioral tendency to form aggregations.

New species of ladybeetles in the genera Scymnus and Mulsantina (Coleoptera: Coccinellidae).
By Chapin, J.B.
Annals of the Entomological Society of America 66: 1071-1073., 1973
New species described:

Scymnus louisianae J. Chapin
Scymnus secures J. Chapin
Scymnus apicanus J. Chapin
Scymnus paracanus J. Chapin
Mulsantina luteodorsa J. Chapin

Chapin, J.B. 1973. New species of ladybeetles in the genera Scymnus and Mulsantina (Coleoptera: Coccinellidae). Annals of the Entomological Society of America 66: 1071-1073.

For more information see full text of:

Chapin, J.B. 1974. The Coccinellidae of Louisiana (Insecta: Coleoptera). Louisiana Agricultural Experiment Station Bulletin 682: 2-87. (1)

The Coccinellidae of Louisiana (Insecta: Coleoptera).
By Chapin, J.B.
Louisiana Agricultural Experiment Station Bulletin 682: 2-87., 1974
Full PDF - I have inserted, as page 2, an undated taxonomic addendum compiled by Chapin.

Chapin, J.B. 1974. The Coccinellidae of Louisiana (Insecta: Coleoptera). Louisiana Agricultural Experiment Station Bulletin 682: 2-87.

Dr. Joan Beggs Chapin (1929-2014)

ABSTRACT:
A taxonomic study of the Coccinellidae of Louisiana was undertaken in 1965 to identify the species occurring in the state. Keys and descriptions are provided for the 6 subfamilies, 11 tribes, 27 genera, and 71 species recorded as occurring in the state. Of the species listed, 38 are reported as new. Distribution records by parish and dates of seasonal occurrences are given.

Notes on Texas Coccinellidae.
By Gaines, J.C.
Bulletin of the Brooklyn Entomological Society 28(5): 211-215., 1933
Full Text

Gaines, J.C. 1933. Notes on Texas Coccinellidae. Bulletin of the Brooklyn Entomological Society 28(5): 211-215.

"The following is a list of Coccinellids collected principally by the writer in the vicinity of College Station and in Hidalgo county, Texas, during 1930, 1931, and 1932. Notes, as far as possible, are given on the habitat, host, and the date that each species was collected. Of the 64 species and varieties listed herein, 19 have not been heretofore reported from this state. The writer is indebted to Dr. E. A. Chapin for determining some of the species of Hyperaspis and Scymnus."

Survey of lady beetles (Coleoptera: Coccinellidae) found on Texas high plains cotton, alfalfa and weeds.
By Carroll, S.C., M.N. Parajulee, and M.D. Arnold.
Proceedings of the World Cotton Conference (September 10–14, 2007), Lubbock, TX., 2007
Full PDF - Manuscript

A study was conducted during the February to November periods of 2004 and 2005 to determine the seasonal presence and reproduction of lady beetle species on cotton, alfalfa, plus over 20 prevalent Texas High Plains weed species. Over the 2-year study period, a total of 221,209 plant sweeps yielded 27,058 lady beetle adults and 3,936 larvae.

The adult specimens represented a total of six different species with the convergent lady beetle, Hippodamia convergens Guerin-Meneville, easily representing over 80% of the overall population.

Determinate evolution in the color-pattern of the lady-beetles.
By Johnson, R.H.
Carnegie Institution of Washington, Publication No. 122. iv + 104 pp., 1910
Full Text

All of the American coccinellids which could be obtained in numbers and which showed a variable color-pattern were studied, viz, Hippodamini, Coccinellae, and Epilachnini, divisions employed by Casey (1899).

Johnson, R.H. 1910. Determinate evolution in the color-pattern of the lady-beetles. Carnegie Institution of Washington, Publication No. 122. iv + 104 pp.

A checklist of the Coccinellidae of Arkansas.
By Rouse, E.P., and J.B. Chapin.
Proceedings of the Arkansas Academy of Science 30: 76-77., 1976
Full PDF

Rouse, E.P., and J.B. Chapin. 1976. A checklist of the Coccinellidae of Arkansas. Proceedings of the Arkansas Academy of Science 30: 76-77.

The list consists of 49 species and includes distribution by county and ecological information if known.

See also:

Rouse, E.P. and J.B. Chapin. 1980. Checklist of the Lady Beetles (Coccinellidae) of Arkansas. Arkansas Academy of Science, Arkansas Biota Survey Checklist No. 26. 2 pp.

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