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Species Heterarthrus nemoratus - Late Birch Leaf Edgeminer

Sawfly - Heterarthrus nemoratus
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon ("Symphyta" - Sawflies, Horntails, and Wood Wasps)
Family Tenthredinidae (Common Sawflies)
Subfamily Heterarthrinae
Tribe Heterarthrini
Genus Heterarthrus
Species nemoratus (Late Birch Leaf Edgeminer)
Other Common Names
Birch Leaf Edge Miner(1)
Explanation of Names
Heterarthrus nemoratus (Fallén 1808)
Size
4.5 mm(1)
Identification
Range
native to Europe & w. Siberia, adventive in NA (across the north)(3)(1)(4)
Food
host: Betula (Birch)(2)
Life Cycle
Parthenogenic; the only birch leafmining sawfly to pupate in the leaf
One generation per year. Overwinters as larvae. Eggs deposited in slits cut in the edges of mature leaves at all levels in the trees. Larvae produce large, blister-like or blotch mines free of frass. Mature larva builds a cocoon or hibernaculum within its mine. The leaf then falls to the ground and the larva stays there throughout the winter.(5)
Remarks
Introduced from Europe early in the 20th century(3); at one time a severe pest of wild birch, now a minor aesthetic pest(1)
In the 1920s/1930s several biocontrol agents were imported, Kratochviliana laricinellae and Phanomeris phyllotomae became established
Internet References
Species pages: Ellis (2007) (Europe)(6) | Edmunds (2007) (UK)(2) | Pitkin et al. (2007-) (UK)(7)