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Genus Sisyra

Spongillafly - Sisyra vicaria Spongillafly - Sisyra vicaria Spongillafly - Sisyra vicaria Spongillafly? - Sisyra vicaria Sisyra sp? - Sisyra vicaria Dark brown lacewing or spongillafly - Sisyra vicaria Dark Brown Spongillafly - Sisyra vicaria My first spongillafly! - Sisyra vicaria
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Neuroptera (Antlions, Lacewings, and Allies)
Suborder Hemerobiiformia (Lacewings and Allies)
Family Sisyridae (Spongilla Lacewings)
Genus Sisyra
Synonyms and other taxonomic changes
Sisyra Burmeister, 1839
Explanation of Names
From the Greek σισύρα (sisýra) 'cloak of goat's hair'
Numbers
3 spp. in our area:(1)
1. Sisyra apicalis: se. US (AL, FL, GA, LA, MS, NC, TX,(2) VA(3)); south to Brazil, West Indies
2. Sisyra nigra (=fuscata): Canada and n. US (BC, ON, QC; AK, IN, MA, ME, MI, MN, NY, WI); w. Palaearctic
3. Sisyra vicaria: s. Canada and transcontinental US (BC, NS, ON, QC; AZ, CT, DC, FL, GA, IL, IN, KS, KY, MA, MD, ME, MI, MN, MO, MS, NC, NY, OK, OR, PA, RI, TN, TX, VA, WA, WI)
Size
3-7 mm(4)
Identification
Differentiated from Climacia by the absence of outer gradates, 3 branches of RS rather than a single branch, and distinctly dark wings.



Wing venation

Keys to larvae and adults in Bowles (2006).(2)
Sisyra apicalis: Body length <5mm, antennae with distinct pale region, forewings streaked
Sisyra nigra: Body length >5mm, antennae uniformly dark brown to black, R2 fork at or apical to Sc / R1 junction
Sisyra vicaria: Body length >5mm, antennae with base (2-3 segments, rarely up to two-thirds) dark brown and remaining segments light brown to yellowish, R2 fork basal to Sc / R1 junction
Range
transcontinental, though significantly more common in the eastern half(5)
See Also
Sympherobius spp. can also have uniformly dark wings, but they can be distinguished from Sisyra by the presence of a series of outer gradates and the branches of R2-R5 connected directly to R1 rather than to a separate Rs vein as in Sisyridae.
Works Cited
1.Species catalog of the Neuroptera, Megaloptera, and Raphidioptera of America North of Mexico
Penny N.D., Adams P.A., Stange L.A. 1997. Proc. Calif. Acad. Sci. 50: 39‒114.
2.Spongillaflies (Neuroptera: Sisyridae) Of North America With A Key To The Larvae And Adults
David E. Bowles. 2006. Zootaxa 1357: 1-19.
3.Annotated Checklist of the Neuropterida of Virginia (Arthropoda: Insecta)
Oliver S. Flint, Jr. 2015. Banisteria, number 45, pages 3-47.
4.The spongilla-flies, with special reference to those of the Western Hemisphere (Sisyridae, Neuroptera)
Parfin, Sophy I. & Gurney, Ashley B. 1956. Proceedings of the United States National Museum, 105(3360): 421-529.
5.Neuropterida Species of the World catalogue (by J.D. Oswald)