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Genus Ululodes - Split-eyed Owlflies

Owlfly - Ululodes floridanus - female Four-spotted Owlfly - Ululodes quadripunctatus - female Four-spotted Owlfly - Ululodes quadripunctatus - male Four-spotted Owlfly - Ululodes quadripunctatus - female Interesting Eyes - Ululodes macleayanus - male Owlfly eggs and hatchllings - Ululodes
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Neuroptera (Antlions, Lacewings, and Allies)
Suborder Myrmeleontiformia (Antlions and Allies)
Family Myrmeleontidae (Antlions including Owlflies)
Subfamily Ascalaphinae (Owlflies)
Genus Ululodes (Split-eyed Owlflies)
Explanation of Names
Ululodes Smith 1900
from Ulula Rambur, 1842 (an owlfly genus-group name) + Greek suffix -ειδής (-odes) 'resembling'
from the Greek ulula ('screech owl')
7 spp. in our area(1) (and comments by J.R. Jones)
1. Ululodes arizonensis Banks, 1907: AZ
2. Ululodes bicolor (Banks, 1895): AZ, CA; records from LA, TX in older literature correspond to separate species (e.g. U. floridanus)
3. Ululodes floridanus (Banks, 1906): AL, FL, GA, LA, NC, SC, TX
4. Ululodes macleayanus (Guilding, 1825): AL, AR, FL, GA, IL, KS, LA, MO, MS, NC, NJ, OK, SC, TX
5. Ululodes mexicanus (McLachlan, 1871): AZ, CA, NM, NV, w. TX, UT
6. Ululodes nigripes Banks, 1943: TX
7. Ululodes quadripunctatus (Burmeister, 1839): widespread, especially eastern (ON; AL, AR, AZ, DC, DE, FL, IA, IL, IN, KS, KY, LA, MD, MI, MO, MS, NC, NE, NJ, NY, OH, OK, PA, SC, TN, TX, VA, WI, WV)
The eyes being divided (or "split") distinguishes this from all other owlfly genera in our area.(2)

In U. quadripunctatus, at least, apparently only male hindwings are stalked(3); female wings are not stalked.

Species identification follows unpublished revision to the genus by Oswald & Jones. Earlier keys such as that in (4) are rendered inadequate under this revision.
fore and hind pterostigmas both dark: U. arizonensis (western, wings distinctively pigmented), U. macleayanus (eastern: widespread), U. mexicanus (western, wings unpigmented), U. nigripes (TX)
fore pterostigmas pale and hind dark: U. bicolor (western), U. floridanus (eastern)
fore and hind pterostigmas both pale: U. quadripunctatus
     U. arizonensis                    U. bicolor                      U. floridanus                U. macleayanus
     U. mexicanus                    U. nigripes                U. quadripunctatus
transcontinental (WI‒ON‒MA to FL‒TX‒CA)
Our only representative of split-eyed antlion (tribes Ululodini and Ascalaphini), that together contain ~340 spp. worldwide