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Genus Ululodes - Split-eyed Owlflies

Owlfly - Ululodes floridanus - female Four-spotted Owlfly - Ululodes quadripunctatus - female Four-spotted Owlfly - Ululodes quadripunctatus - male Four-spotted Owlfly - Ululodes quadripunctatus - female Interesting Eyes - Ululodes macleayanus - male Owlfly eggs and hatchllings - Ululodes
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Neuroptera (Antlions, Lacewings, and Allies)
Suborder Myrmeleontiformia (Antlions and Allies)
Family Myrmeleontidae (Antlions including Owlflies)
Subfamily Ascalaphinae (Owlflies)
Genus Ululodes (Split-eyed Owlflies)
Other Common Names
Split‐eyed Owlflies (note: this is the only such genus found in the area, but the trait is globally shared between the tribes of Ascalaphini and Ululodini)
Explanation of Names
Ululodes Currie in Smith 1899
from Ulula Rambur, 1842 (an owlfly genus-group name) + Greek suffix -ειδής (-odes) 'resembling'
from the Greek ulula ('screech owl')
Numbers
7 spp. in our area(1) (and comments by J.R. Jones)
1. Ululodes arizonensis Banks, 1907: restricted southwestern (AZ)
2. Ululodes bicolor (Banks, 1895): AZ, CA; records from LA, TX in older literature correspond to separate species (e.g. U. floridanus)
3. Ululodes floridanus (Banks, 1906): AL, FL, GA, LA, NC, SC, TX
4. Ululodes macleayanus (Guilding, 1825): AL, AR, FL, GA, IL, KS, LA, MO, MS, NC, NJ, OK, SC, TX
5. Ululodes mexicanus (McLachlan, 1871): AZ, CA, NM, NV, w. TX, UT
6. Ululodes nigripes Banks, 1943: TX
7. Ululodes quadripunctatus (Burmeister, 1839): widespread, especially eastern (Canada: ON; United States: AL, AR, AZ, DC, DE, FL, IA, IL, IN, KS, KY, LA, MD, MI, MO, MS, NC, NE, NJ, NY, OH, OK, PA, SC, TN, TX, VA, WI, WV)
Size
4-5 cm
Identification
The eyes being divided (or "split") distinguishes this from all other owlfly genera in our area.(2)

In U. quadripunctatus, at least, apparently only male hindwings are stalked(3); female wings are not stalked.


Overview of Species
Species with fore and hind pterostigmas both dark: U. arizonensis (western, wings distinctively pigmented), U. macleayanus (eastern: widespread), U. mexicanus (western, wings unpigmented), U. nigripes (eastern: TX only)
Species with fore pterostigmas pale and hind dark: U. bicolor (western), U. floridanus (eastern)
Species with fore and hind pterostigmas both pale: U. quadripunctatus
                 
     U. arizonensis                    U. bicolor                      U. floridanus                U. macleayanus

           
     U. mexicanus                    U. nigripes                U. quadripunctatus
Remarks
Our only representative of split-eyed antlion (tribes Ululodini and Ascalaphini), that together contain ~340 spp. worldwide