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For Insects, Spiders & Their Kin
For the United States & Canada

Tribe Pentodontini


Collagenus dasysternus ... with a key to the New World genera of Pentodontini
By Brett Ratcliffe and Martin Hardy
The Coleopterists Bulletin, Vol. 59, pp.143-150, 2005
Brett Ratcliffe and Martin Hardy, Collagenus dasysternus, a new genus and species of Dynastinae from eastern Venezuela with a key to the New World genera of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 59(1): 143-150, 2005.

From BG's Pentodontini Info tab, "One error noticed; couplet 11- range of genus Aphonus should be 'United States east of the Rocky Mountains'".

Full text

A Review of the North American Genus Aphonus Leconte (Coleoptera: Scarabaeidae: Dynastinae)
By B. D. Gill and H. F. Howden
The Coleopterists Bulletin 39(2): 119-129, 1985
Here is a link to the first page.

The Dynastine Scarab Beetles of the USA and Canada (Coleoptera: Scarabaeidae: Dynastinae)
By Brett C. Ratcliffe & Ronald D. Cave
University of Nebraska State Museum, Vol 30, 298 pp, 2017
The 61 species and two subspecies of dynastine scarab beetles that occur in the United States of America (including Guam) and Canada are reviewed. Discussions of historical collecting, people, climate, vegetation, and habitats are presented. Keys to all tribes, genera, and species in the study area are given. Descriptions, recorded geographic localities and temporal distributions, diagnoses, notes on natural history, illustrations, and distribution maps are provided for all species. Also included are synopses of the higher-level taxa of the subfamily in the region, a glossary, and a species checklist.

Review of the genus Orizabus Fairmaire in the United States of America (Coleoptera: Scarabaeidae: Dynastinae)
By W.B. Warner
Insecta Mundi 0174: 1-42, 2011

Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae) new classification and taxonomic revision
By López-García, M.M. and C. Deloya
Zootaxa 5211(1): 1-119., 2022
Abstract and References - Zootaxa

López-García, M.M. and C. Deloya. (2022) Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision. Zootaxa 5211(1): 1-119.

Abstract

The phylogeny of Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) was explored by parsimony using morphological characters to approach its taxonomic and nomenclature problems. The genus was not supported as a monophyletic group, and the recovered lineages did not match at all any of the previously known classifications. Three different lineages and six monophyletic groups were recovered, which could be treated as independent genera. However, the classification proposed here is as conservative as possible concerning to the creation of many new names and generic taxa.

Scarab beetles of the genus Bothynus in the United States (Coleoptera: Scarabaeidae)
By O. Cartwright
Proceedings of the United States National Museum, 108(3409): 515-541, 1959
Online here.

Monograph of the genus Tomarus (=Bothynus, Ligyrus)

Taxonomic review of the North American dung beetle genus Melanocanthon Halffter, 1958
By Edmonds, W.D.
Insecta Mundi, 2023
Edmonds, W.D. 2023. Taxonomic review of the North American dung beetle genus Melanocanthon Halffter, 1958 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Insecta Mundi 1014: 1-28.

New country and state Records of Scarabaeidae (Coleoptera) in the USA.
By Warner, W.B.
The Coleopterists Bulletin 76(1): 128-129., 2022
BioOne

Warner, W.B. (2022). New country and state Records of Scarabaeidae (Coleoptera) in the USA. The Coleopterists Bulletin 76(1): 128-129.

This brief paper describes new Arizona state distribution records for Haroldiellus sallei (Harold) (Scarabaeidae: Aphodiinae: Aphodiini) and Phyllophaga crinita (Burmeister) (Scarabaeidae: Melolonthinae: Rhizotrogini), as well as new USA country records (Arizona and Texas, respectively) for Haroldiataenius limbatus (Bates) (Scarabaeidae: Aphodiinae: Eupariini) and Amithao erythropus (Burmeister) (Scarabaeidae: Cetoniinae: Gymnetini). Vouchers of all the species are deposited in the Hasbrouck Insect Collection at Arizona State University, Tempe, Arizona, USA.