Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
Moths Butterflies Flies Caterpillars Flies Dragonflies Flies Mantids Cockroaches Bees and Wasps Walkingsticks Earwigs Ants Termites Hoppers and Kin Hoppers and Kin Beetles True Bugs Fleas Grasshoppers and Kin Ticks Spiders Scorpions Centipedes Millipedes

Fall Fund Drive

TaxonomyBrowse
Info
ImagesLinksBooksData

Tribe Eupariini

Dung beetle? - Ataenius strigatus Coleoptera - Ataenius platensis Euparixia duncani Brown - Euparixia duncani Aphodiine Dung Beetle - Ataenius strigatus Dung Beetle? - Ataenius platensis Ataenius wenzelii Horn - Ataenius wenzelii Ataenius sp.? - Ataenius strigatus Aphodius granarius? - Ataenius
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Scarabaeidae (Scarab Beetles)
Subfamily Aphodiinae (Aphodiine Dung Beetles)
Tribe Eupariini
Synonyms and other taxonomic changes
Lomanoxiini Stebnicka 1999
Explanation of Names
Eupariini Schmidt 1910
Numbers
~70 spp. in 10 genera in our area, >340 spp. in 31 genera in the New World, ~570 spp. in ~45 genera worldwide(1)
Tribe Eupariini [updated from(1)]
Aphotaenius carolinus (Van Dyke); eastern US
Ataeniopsis duncani (Cartwright); Central and SW-USA
Ataeniopsis edistoi Cartwright; SC, FL
Ataeniopsis figurator (Harold); SE-USA, Mexico
Ataeniopsis parkeri (Cartwright); SW-USA, Mexico
Ataeniopsis rugopygus (Cartwright); Central and SW-USA
Ataeniopsis saxatilis (Cartwright); GA, SC, NC
Genus Ataenius
Ataenius aequalis Harold; N-Argentina to S- US, and West Indies
=Ataenius insulicola Chapin
=Ataenius titschacki Balthasar
Ataenius alternatus (Melsheimer); SE-US, Mexico
Ataenius apicalis Hinton; Honduras, Mexico, E-USA
Ataenius abditus (Haldeman); widespread New World
Ataenius barberi Cartwright; TX,AZ-USA
Ataenius brevicollis (Wollaston); SE-USA, West Indies, Madeira Is., Canary Is., Portugal
=Pleurophorus chopardi Paulian
=Ataenius frankenbergeri Balthasar
=Ataenius lherminiari Paulian
=Auperia sulcatula Chevrolat
Ataenius brevis Fall; E-USA - Type
Ataenius californicus Horn; SW-USA - Type
Ataenius carinator Harold; Mexico to S. America, FL-USA (introduced), West Indies
=Ataenius vincentiae Arrow
Ataenius cognatus (LeConte); SW-USA, Mexico
Ataenius confertus Fall; SW-USA, Baja California
=Ataenius cribratus VanDyke
Ataenius cylindrus Horn; SE-USA - Type
=Ataenius cylindricus Horn in Schmidt (misspelling)
=Ataenius horni Harold
Ataenius desertus Horn; SW-USA, Mexico - Type
Ataenius ecruensis Stebnicka and Lago; E-USA
Ataenius erratus Fall; SE-USA - Type
Ataenius exiguous Brown; SE-USA
Ataenius fattigi Cartwright; E-USA
Ataenius glaseri Cartwright; E-USA
Ataenius gracilis (Melsheimer); US to South America, West Indies, Micronesia, Pacific Islands
=Saprosites nocturnus Nomura
Ataenius heinekeni (Wollaston); SE-US to Central America, Cuba
=Auperia rhyticephalus Chevrolat
=Ataenius floridanus Brown
=Ataenius solitarius Blatchley
=Ataenius vexator Harold
Ataenius hesperius Cartwright; W-USA
Ataenius hirsutus Horn; SW-USA, Mexico - Type
Ataenius imbricatus (Melsheimer); Central and E-USA to Argentina, W.Indies
=Ataenius sordidus Harold
Ataenius inquisitus Horn; SW-USA - Type
Ataenius insculptus Horn; SE-USA - Type
=Ataenius sculptilis LeConte 1878 [not Ataenius sculptilis Harold 1868]
Ataenius languidus Schmidt; S-USA to Venezuela, West Indies
=Ataenius linelli Cartwright
Ataenius lobatus Horn; SW-USA, Baja California-Mexico - Type
=Ataenius laeviventris Horn
Ataenius miamii Cartwright; E-USA, Bahamas
Ataenius oklahomensis Brown; S. Central USA (AK, LA, OK, TX)
Ataenius opatrinus Harold; Central and S. America [FL-USA?]
=Ataenius narialis Petrovitz
=Ataenius paraperforatus Deloya and Ibanez-Bernal
=Ataenius woodruffi Cartwright
Ataenius ovatulus Horn; SE-USA (MD-IL and south) - Type
=Ataenius lecontei Harold
Ataenius peregrinator Harold; Florida to AZ; Indonesia, Philippines, Micronesia, Vietnam, Colombia and Ecuador.
=Ataenius jalapensis Bates (See Galante et al. 2003)
=Ataenius polyglyptus var. jalapensis
=Ataenius polyglyptus var. hieronymi
=Ataenius nunenmacheri Cartwright
=Ataenius stroheckeri Cartwright
=Ataenius sumatrensis Balthasar
Ataenius picinus Harold; nearly cosmopolitan (Americas, Australia, Fiji, Vanuatu)
=Ataenius alegrus Balthasar
=Ataenius boucomontii Paulian
=Ataenius darlingtoni Hinton
=Ataenius duplopunctatus Lea
=Ataenius paracognatus Balthasar
=Ataenius queirosii Paulian
=Saprosites rugosus Richards
=Ataenius salutator Fall
Ataenius platensis (Blanchard); Argentina to S-USA, W. Indies
=Ataenius anticus Fall - Type
=Ataenius degallieri Chalumeau
=Ataenius granchacoensis Balthasar
=Ataenius heyrovskyi Balthasar
=Ataenius histrionicus Balthasar
=Ataenius integer Harold
Ataenius pseudocarinator Balthasar; US, French Guiana
=Ataenius vandykei Cartwright
Ataenius robustus Horn; Central-US - Type
Ataenius scabrelloides Petrovitz; Florida; Costa Rica to Bolivia to Brazil
=Ataenius superficialis Cartwright
Ataenius scabrellus Schmidt; Venezuela, West Indies to south Florida-USA
=Ataenius havanensis Balthasar
=Ataenius fleutiauxi Paulian
Ataenius sculptor Harold; SW-US to Colombia
=Ataenius oblongus Horn - Type
Ataenius setiger Bates; Mexico, SW-US
=Ataenius pseudohirsutus Cartwright
Ataenius spretulus (Haldeman); SE-Canada, US
=Ataenius consors Fall 1930 [not Blackburn 1904] - Type
=Ataenius falli Hinton
Ataenius stephani Cartwright; AZ-USA, Mexico
Ataenius strigatus (Say); SE-Canada, USA, Mexico
Ataenius texanus Harold; Central USA, Mexico, Central America and West Indies
=Ataenius punctifrons Cartwright
Ataenius utahensis Cartwright; Utah-US
Ataenius waltherhorni Balthasar; E-US, Cuba [was Pseudatanius sp. in Cartwright]
Ataenius wenzelii Horn; SE-USA - Type
=Ataenius ludovicianus Fall - Type
=Ataenius rudellus Fall - Type
Odontolytes denominatus (Chevrolat); USA-FL, Cuba, Mexico and Central America to South America
=Auperia stercorator Chevrolat 1864 [not Ataenius stercorator of Fabricius 1775]
=Ataenius arator Harold
=Ataenius stercorator of Harold 1870 [not Fabricius 1775, not Chevrolat 1864]
=Ataenius euglyptus Bates
=Ataenius denominatus in Schmidt (as synonym of Ataenius stercorator Fabricius)
=Ataenius brevinotus Chapin - Type
=Ataenius benjaminbanderai Islas
=Ataenius sciurus Cartwright, Woodruff 1973
=Phalangochaeta grandis Petrovitz
Genus Euparia
Euparia castanea LePeletier and Serville; se US
Genus Euparixia
Euparixia duncani Brown; S.AZ-USA, Mexico
Euparixia moseri Woodruff and Cartwright; E.TX, LA-USA
Haroldiataenius buvexus Stebnicka & Skelley; Big Bend region Texas
Haroldiataenius convexus (Robinson); Southern coastal Texas
Haroldiataenius griffini (Cartwright); Texas
Haroldiataenius lucanus (Horn); Arizona, Baja California, Mexico - Type
Haroldiataenius saramari (Cartwright); Florida
Haroldiataenius semipilosus (Van Dyke); Texas and Arizona, Mexico
Martineziana dutertrei (Chalumeau); Southern US, South America
Parataenius simulator (Harold); Southern US, South America
Pseudataenius contortus Cartwright; Eastern and southeastern USA, Alabama, Delaware, Florida, Louisiana, Maryland, Mississippi, New Jersey, South Carolina, Virginia
Pseudataenius socialis (Horn); USA, Kansas, Louisiana, Nebraska, Oklahoma, Texas - Type

Recorded from Hawaii; Saprosites pygmaeus Harold, Ataenius pacificus (Sharp)
Range
Worldwide(1)
Print References
Stebnicka Z. 2001. The New World species of Ataenius, Harold, 1867.I. Revision of the A. crenator-group, A. nugator-group and A. perforatus-group (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta zoologica cracoviensia 44(3): 253-283.
Stebnicka Z. 2002. The New World Eupariini: Revision of the genus Auperia Chevrolat, 1864 (=Phalangochaeta Martinez, 1952) (Coleoptera: Scarabaeidae: Aphodiinae). Revue suisse de Zoologie 109(4): 741-775.
Stebnicka Z. 2002. The New World species of Ataenius Harold, 1867. II. Revision of the West Indian A. terminalis-group. (Col. Scarabaeidae: Eupariini). Acta zoologica cracoviensia, 45(3): 259-281.
Stebnicka Z. 2003. Revision and hypothetical phylogenetic analysis of the species of the New World genus Ataeniopsis (Coleoptera: Aphodiinae: Eupariini). European Journal of Entomology 100: 101-113.
Stebnicka Z. 2003. The New World species of Ataenius Harold, 1867. III. Revision of the A. imbricatus-group (Coleoptera: Scarabaeidae: Eupariini). Acta zoologica cracoviensia 46(3): 219-249.
Stebnicka Z. 2003. Revision and hypothetical phylogenetic analysis of the species of the New World genus Ataeniopsis (Coleoptera: Aphodiinae: Eupariini). European Journal of Entomology 100: 101-113.
Stebnicka Z. 2004. The New World species of Ataenius Harold, 1867.IV. Revision of the A. strigicauda-group (Coleoptera: Scarabaeidae: Eupariini). Acta zoologica cracoviensia 47: 211-228.
Stebnicka Z. 2005. The New World species of Ataenius Harold, 1867.VI. Revision of the A. aequalis-platensis-group (Coleoptera: Scarabaeidae:Aphodiinae: Eupariini). Acta zoologica cracoviensia 48B(1-2): 99-138.
Stebnicka Z. & P.E. Skelley. 2005. Review of some New World aphodiine genera and descriptions of new species (Coleoptera: Scarabaeidae: Aphodiinae). Acta zoologica cracoviensia 48B(1-2): 23-42.
Stebnicka Z. & P.K. Lago. 2005. The New World species of Ataenius Harold, 1867.VI. Revision of the A. strigatus-group (Coleoptera: Scarabaeidae:Aphodiinae: Eupariini). Insecta Mundi, 19:55-83. Download PDF
Stebnicka Z. 2006. The New World species of Ataenius Harold, 1867.VI. Revision of the A. complicatus-group (Coleoptera: Aphodiinae: Eupariini). Acta zoologica cracoviensia 49B(1-2): 89-114.
Stebnicka Z. 2006. Aegialiinae, Eremazinae, Didactyliini, Eupariini, Thinorycterini (102-105; 142-144, 179). In: Catalogue of Palaearctic Coleoptera, vol. 3: Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. (Eds. Loebl & Smetana). Apollo Books, Stenstrup, 690 pp.
Stebnicka Z. 2007. The New World species of Ataenius Harold, 1867. VIII. Revision of the A. scutellaris group and diagnosis of the A. texanus-carinator group with descriptions of new species (Coleoptera: Scarabaeidae: Aphodiinae: Eupariini). Acta zoologica cracoviensia, 50B(2): 45-81.
Stebnicka Z. & P.E. Skelley. 2009. A revision of the genus Haroldiataenius Chalumeau (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi, 62: 1-16.
Stebnicka Z. & P.E. Skelley. 2009. Taxonomic redefinition of the genera Parataenius Balthasar and Pseudataenius Brown, with descriptions of three new species (Scarabaeidae: Aphodiinae: Eupariini). Insecta Mundi, 66: 1-18.
Krell, F.-T., G. Dellacasa & M. Dellacasa. 2011. Auperia Jacquelin-Duval is a junior synonym of Euparia Le Peletier de Saint-Fargeau and Serville and Odontolytes Kozhantshikov is the valid name for Auperia sensu Stebnicka (Coleoptera: Scarabaeidae: Aphodiinae). Coleop. Bull. 65(2):173-176
Internet References