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Hawkweed mountaineer fly - Tmemophlebia - female

Hawkweed mountaineer fly - Tmemophlebia - Female
Parker River NWR, Newbury, Essex County, Massachusetts, USA
July 25, 2010
Size: 2.5 mm
Female Apolysis sigma?

Images of this individual: tag all
Hawkweed mountaineer fly - Tmemophlebia - female Head - Tmemophlebia - female In better color - Tmemophlebia

I don't have the revision at hand, but I think T. coquilletti is the only species likely in Massachusetts.

Moved from Phthiriinae.

According to the world catalog linked from genus page, T. coquilletti is known from southern Canada and the United States from the Rocky Mountains to the Atlantic, plus California. T. aldrichi is known from Idaho south in the west but only from Georgia in the east.

The world catalog was last updated in 2003, while the genus was revised in 2004 (Hall, J.C., and Evenhuis, N.L. 2004. Bombyliidae. In Flies of the Nearctic Region. Edited by G.C.D. Griffiths. E. Schweizerbart, Stuttgart. Vol. V, pt. 13, no. 7, pp 657–716.), so those distributions aren't entirely accurate and a number of new species have been described that aren't in the catalog.

You're both right...only 1 species recorded from MA
I just carefully scanned the distribution records given for each of the 9 species of Tmemophlebia listed in Hall & Evenhuis(1) as occurring north of Mexico. Turns out T. coquilletti is the only species recorded from Massachusetts.

The very detailed descriptions of male and female of that species (with plenty of non-genitalic characters) appear on pg. 675. At this point, I don't feel I have the expertise with this group to evaluate the fit of the images here with that description...but, given range considerations, it's a natural candidate.

Moved from Bee Flies. First record for the subfamily on BugGuide from northeast USA.

I think you're right.
But have you checked subfamily Phthiriinae as well? (I've tripped over these a couple times.)

This is one of those genera related to Phthiria. Apolysis has one fewer posterior cell. Look at the discal cell: In Phthiriinae, there is a vein running from the center of the cell's posterior margin to the wing edge; in Apolysis, this vein is lacking. I believe the two belong in the same subfamily, which is where Yeates put them. The antennal form clearly unites them.

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