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Subfamily Agathidinae

red & black Wasp with dark smokey wings and long ovipositor Cremnops ashmeadi - female unknown - red abdomen - Alabagrus texanus - female Wasp oviposting in Burdock (Arctium lappa) flower. - Agathis malvacearum - female Braconid June 10 Earinus - Earinus limitaris - male Ichneumon - Genus Arenetra? - Earinus - male Braconid/ Agathidinae/ Aleiodes? - female
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon ("Parasitica" - Parasitoid Wasps)
Superfamily Ichneumonoidea (Braconid and Ichneumonid Wasps)
Family Braconidae (Braconid Wasps)
Subfamily Agathidinae
Synonyms and other taxonomic changes
4 tribes recognized in (1), 7 tribes more recently(2)
Explanation of Names
Agathidinae Haliday 1833
Numbers
~100 species(3), 16 genera (+1 undescribed) in our area(4); ~2,000 spp. in ~40 genera worldwide.
Tribe Agathidini: Aerophilus (35 spp.), Agathis, Alabagrus (6 spp.), Aphelagathis (6 spp.) Neothlipsis, Plesiocoelus, Pneumagathis
Tribe Agathirsini: Agathirsia, Crassomicrodus, Gelastagathis
Tribe Cremnoptini: Cremnops, Zacremnops
Tribe Disophrini: Zelomorpha (3 spp.)
Tribe Earinini: Earinus (5 spp.)
Tribe Lytopylini: Lytopylus (2 spp.)
Tribe Mesocoelini: Therophilus, undescribed genus (=annulipes-group)
Identification
Keys to genera in (3)(5)(4)
Fore wing M+CU not tubular in basal 1/3 or more
Fore wing 3RSb terminating at wing margin near stigma and distant from wing apex
Range
Worldwide
Habitat
Most terrestrial habitats. Agathis and Earinus are more species-rich in temperate regions. In Agathis and Crassomicrodus, some species tend toward desert habitat. Other genera most diverse in humid tropics.(6)
Food
hosts: Lepidoptera larvae
Life Cycle
Remarks
species useful for pest control in our area include Agathis pumila against larch casebearer; A. stigma and Cremnops laricella against sugarcane borer; & Bassus unicoloratus against potato tuberworm.
"There are three main Agathidine genera with the 'long face' (Agathus, Bassus, and Cremnops). Agathus is almost always entirely black, whereas Bassus and Cremnops are red & black. Cremnops has a carina extending from the lateral margin of the antenna to the lateral ocelli. The carina is absent in Bassus." (Comment by M.J. Sharkey, 2007).(7)
Bassus has been split into several genera; according to the current concept, Bassus do not occur in our area.
Works Cited
1.Revision of the Agathidinae (Hymenoptera: Braconidae) with comparisons of static and dynamic alignments
Sharkey M.J., Laurenne N.M., Sharanowski B., Quicke D.L.J., Murray D. 2006. Cladistics 22: 546–567.
2.Phylogeny of the Agathidinae (Hymenoptera: Braconidae) with a Revised Tribal Classification and the Description of a New Genus
Michael J Sharkey & Eric G Chapman. 2017. Entomological Society of Washington 119(sp1):823-842.
3.Synopsis of the Agathidiinae (Hymenopetra: Braconidae) of America north of Mexico
Sharkey M.J. 2004. Proc. Rus. Ent. Soc. 75: 134‒152.
4.Description of Chilearinus Sharkey gen. nov. and status of Nearctic Earinus Wesmael, 1837 with the description of new species
Michael J. Sharkey, Austin Baker, Ramya Manjunath, & Paul D. N. Hebert. 2022. ZooKeys 1099: 57-86.
5.Two new genera of Agathidinae (Hymenoptera: Braconidae) with a key to the genera of the New World
M. Sharkey. 2006. Zootaxa 1185: 37–51.
6.Manual of the New World Genera of the Family Braconidae (Hymenoptera)
Wharton, R.A., P.M. Marsh, M.J. Sharkey (Eds). 1997. International Society of Hymenopterists.
7.Bassus erythrogaster? (Braconidae) in the Colorado Rockies