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Subfamily Agathidinae

Wasp for ID. Spider wasp, perhaps? red & black Wasp with dark smokey wings and long ovipositor wasp - Genus-novum annulipes - female Wasp - Earinus limitaris - male Cremnops sp. - Cremnops - female Agathirsia? - Agathirsia - female Cremnops sp. - Cremnops Wasp - Earinus
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hymenoptera (Ants, Bees, Wasps and Sawflies)
No Taxon ("Parasitica" - Parasitoid Wasps)
Superfamily Ichneumonoidea (Braconid and Ichneumonid Wasps)
Family Braconidae (Braconid Wasps)
Subfamily Agathidinae
Synonyms and other taxonomic changes
Higher taxonomy revised by Sharkey et al. (2006) to include 4 tribes.(1)
Higher taxonomy further revised by Sharkey & Chapman to include 7 tribes.(2)
Explanation of Names
Agathidinae Haliday, 1833
~100(3) spp. 15 (+ 1 undescribed) genera in our area,(4) ~2,000 spp. in ~40 genera worldwide

1. Tribe Agathidini: Aerophilus (35 spp.), Agathis, Alabagrus (6 spp.), Neothlipsis, Plesiocoelus, Pneumagathis
2. Tribe Agathirsini: Agathirsia, Crassomicrodus, Gelastagathis
3. Tribe Cremnoptini: Cremnops, Zacremnops
4. Tribe Disophrini: Zelomorpha (3 spp.)
5. Tribe Earinini: Earinus (5 spp.)
6. Tribe Lytopylini: Lytopylus (2 spp.)
7. Tribe Mesocoelini: Therophilus
undescribed gen. nov. (annulipes-group)
Keys to genera in Sharkey (2004),(3) Sharkey (2006),(5) and Sharkey et al. (2022).(4)
Fore wing M+CU not tubular in basal 1/3 or more
Fore wing 3RSb terminating at wing margin near stigma and distant from wing apex
Most terrestrial habitats. Genera Agathis and Earinus are more species-rich in temperate regions. In Agathis and Crassomicrodus, some species tend toward desert habitat. Other genera most diverse in humid tropics.(6)
hosts: Lepidoptera larvae
Life Cycle
Among the many species useful for insect pest control in the Nearctic are: Agathis pumila against larch casebearer; A. stigma and Cremnops laricella against sugarcane borer; & Bassus unicoloratus against potato tuberworm.

"There are three main Agathidine genera with the 'long face' (Agathus, Bassus, and Cremnops). Agathus is almost always entirely black, whereas Bassus and Cremnops are red & black. Cremnops has a carina extending from the lateral margin of the antenna to the lateral ocelli. The carina is absent in Bassus." (Comment by Dr. Michael J. Sharkey, 2007).(7)
Editor's note: further revision has split the former understanding of Bassus into additional genera such that no Bassus occur in our area.
Works Cited
1.Revision of the Agathidinae (Hymenoptera: Braconidae) with comparisons of static and dynamic alignments
Sharkey M.J., Laurenne N.M., Sharanowski B., Quicke D.L.J., Murray D. 2006. Cladistics 22: 546–567.
2.Phylogeny of the Agathidinae (Hymenoptera: Braconidae) with a Revised Tribal Classification and the Description of a New Genus
Michael J Sharkey & Eric G Chapman. 2017. Entomological Society of Washington 119(sp1):823-842.
3.Synopsis of the Agathidiinae (Hymenopetra: Braconidae) of America north of Mexico
M.J. Sharkey. 2004. Proc. Russian Entomol. Soc. 75(1): 134-152.
4.Description of Chilearinus Sharkey gen. nov. and status of Nearctic Earinus Wesmael, 1837 with the description of new species
Michael J. Sharkey, Austin Baker, Ramya Manjunath, & Paul D. N. Hebert. 2022. ZooKeys 1099: 57-86.
5.Two new genera of Agathidinae (Hymenoptera: Braconidae) with a key to the genera of the New World
M. Sharkey. 2006. Zootaxa 1185: 37–51.
6.Manual of the New World Genera of the Family Braconidae (Hymenoptera)
Wharton, R.A., P.M. Marsh, M.J. Sharkey (Eds). 1997. International Society of Hymenopterists.
7.Bassus erythrogaster? (Braconidae) in the Colorado Rockies