33 spp. (incl. one adventive) in 6 subgenera in our area, ~50 spp. in 9 subgenera total(1)
best separated from Harpalus by having the mentum and submentum partially (usually over the lateral lobes) or entirely fused and spongy adhesive setae underneath some of the pro- and mesotarsomeres of the males(2)
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