Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Genus Amphicosmus

Amphicosmus arizonicus? - Amphicosmus arizonicus Amphicosmus arizonicus Amphicosmus arizonicus Fly - Amphicosmus arizonensis Orthorrhapha - Amphicosmus arizonicus Orthorrhapha - Amphicosmus arizonicus Orthorrhapha - Amphicosmus arizonicus Bombyliidae - Amphicosmus elegans - female
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Diptera (Flies)
No Taxon (Orthorrhapha)
Superfamily Asiloidea
Family Bombyliidae (Bee Flies)
Subfamily Tomomyzinae
Genus Amphicosmus
Explanation of Names
Amphicosmus Coquillett 1891
Numbers
5 spp., all in our area(1)
  Subgenus Amphicosmus Coquilett, 1891 (see also Hull, 1973)
      arizonensis Johnson & Johnson, 1959
      arizonicus Hall, 1975
      elegans Coquillett, 1891
      vanduzeei Cole, 1922
  Subgenus Glycophorba Hull, 1973
      cincturus Williston, 1901
Identification
• Genus Amphicosmus is distinctive among members of subfamily Tomomyzinae in having a cross-vein joining R2+3 and R4, which gives rise to 3 submarginal cells:

• There are 5 extant species, they can be "partly" separated as follows:
    A. arizonicus is distinguished by its conspicuous dark wing markings (the other 4 Amphicosmus species have entirely hyaline wings):
       
    A. elegans and A. arizonensis have proximal segments of the abdomen deep red (the other 3 Amphicosmus species have ground color of abdomen black):
       
    A partial key treating A. vanduzeei, A. elegans, and A. cincturus appears in Hall(1957)
• A very detailed description of genus Amphicosmus is given in the 1973 book "Bee Flies of the World" by Hull(2).
Range
AZ-CA(1)
      arizonensis : Arizona
      arizonicus : Arizona
      cincturus : Arizona & California
      elegans : California
      vanduzeei : California
See Also
In genus Amphicosmus there is a cross-vein joining R2+3 and R4, thus forming 3 submarginal cells:

Our other two genera in the subfamily Tomomyzinae, Paracosmus and Metacosmus, can be similar looking to Amphicosmus, but they have no cross-vein joining R2+3 and R4, and thus those genera have only two submarginal cells.
Print References
Hall, J. C. (1957) Notes and descriptions of new California Bombyliidae (Diptera). Pan-Pac. Entomologist 33: 141–48. (Full Text)
Melander A.L. (1950) Taxonomic notes on some smaller Bombyliidae (Diptera). Pan-Pacific Entomologist 26: 139-156. (Full Text)
Internet References
Specimen Records for Amphicosmus from the California Academy of Sciences
• Figures from Hull(2)(1973) for Amphicosmus:
      antenna of A. cincturus (Fig. 110) and A. elegans (Fig. 114) here;
      wing venation of A. cincturusFig. 361;
      frontal view of head for A. cincturusFig. 608;
      profile view of head for A. cincturusFig. 618;
• Figures from Curran(3)(1933) for Amphicosmus cincturus:
      head profile — Fig. 50
      dorsal view of body & wing venation — Fig. 53