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Genus Anillinus

Anillinus stephani Sokolov & Carlton - Anillinus stephani - female Anillinus cherokee Sokolov & Carlton - Anillinus cherokee Anillinus barberi - male Anillinus folkertsi - male Anillinus daggyi - male Anillinus moseleyae - male Anillinus gimmeli - male Anillinus folkertsioides - male
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Trechinae
Tribe Bembidiini
Subtribe Anillina
Genus Anillinus
Synonyms and other taxonomic changes
Micranillodes Jeannel 1963(1)
Explanation of Names
Anillinus Casey 1918
Numbers
64 described species (as of June 2021), all in our area(1), (2), (3), (4), (5), (6),(7),(8).

2 species described from Brazil were revealed to actually be mislabeled specimens collected in Cloudland Canyon State Park, Georgia (9).
Identification
Accurate identification to species level requires examination of male genitalia.
There is no complete key to species.
Partial regional keys:

key to the Texas species(3)
key to eastern species known as of 2004(10), new eastern species described in 2007(11), 2008(12), 2010(13), 2011(14), 2014(5)
key to species known from AR, MO, and OK as of 2008(15), new AR species described 2017(4)
key to the Alabama species(6) (new species described in(7))
Range
Most diverse in the southern Appalachians(16)(1)(10); north to MD, west to IN, TX, AR, MO(17), south to LA(7), AL, FL(8).
Habitat
in leaf litter or under large embedded stones on mountain slopes or in ravines in forested areas; some in caves(16), deep sand(8)
Season
Year-round.
Remarks
Completely eyeless.

Over 40 undescribed species are known (C. Harden, personal observation).
Works Cited
1.Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico
Bousquet Y. 2012. ZooKeys 245: 1–1722.
2.Five new species of Anillinus Casey from Alabama with a key to the Alabama species (Carabidae: Trechinae: Bembidiini)
Sokolov, I.M. 2012. Annals of Carnegie Museum 81:61-71.
3.Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae...)
Sokolov I.M., Reddell J.R, Kavanaugh D.H. 2014. ZooKeys 417:71-101.
4.Anillinus alleni Sokolov and Carlton (Coleoptera: Carabidae: Trechinae: Bembidiini), a New Species from the Ozark [...]
Igor Sokolov, C. E. Carlton, L. E. Watrous, and H. W. Robison. 2017. Coleopterists Bulletin.
5.A New Species of Anillinus Casey from the Oak Ridge Area West of the Appalachian Mountains in Tennessee [...]
Igor Sokolov. 2014. Annals of Carnegie Museum.
6.Four new species of the genus Anillinus Casey (Coleoptera, Carabidae, Anillini) from Alabama, U.S.A., with a revised key [...]
Igor M. Sokolov. 2020. Zootaxa 4808 (3): 547-559.
7.Two new species of the genus Anillinus Casey (Coleoptera, Carabidae, Anillini) from the southern United States
Igor Sokolov. 2021. Zookeys 1016: 63–76.
8.A new subterranean species of Anillinus Casey (Carabidae, Trechinae, Anillini) from Florida
Sokolov & Schnepp. 2021. Subterranean Biology.
9.Species of Anillinus Casey (Coleoptera: Carabidae: Trechinae) Described from Brazil and their Relation to North American [...]
Igor Sokolov and Chris Carlton. 2012. Coleopterists Bulletin.
10.Review of Anillinus, with descriptions of 17 new species and a key to soil and litter species (Coleoptera: Carabidae: Trechinae:
Sokolov I.M., Carlton C., Cornell J.F. 2004. Coleopterists bulletin 58: 185-233.
11.New species of Anillinus Casey (Carabidae : Trechinae : Bembidiini) from Great Smoky Mountains National Park, USA and [...]
Igor M. Sokolov, Yuliya Y. Sokolova, and Christopher E. Carlton. 2007. Zootaxa.
12.Two new species of blind, forest litter-inhabiting ground beetles from the subtribe Anillina [...]
Igor Sokolov and Chris Carlton. 2008. Zootaxa.
13.New Species Of Anillinus Casey (Carabidae: Trechinae: Bembidiini) From The Southern Appalachians And Phylogeography Of The [...]
Igor Sokolov and Chris Carlton. 2010. Zootaxa.
14.Five new species of Anillinus Casey from the Southern Appalachian Mountains and the Piedmont Plateau of eastern U.S.A.
Igor M. Sokolov. 2011. Insecta Mundi.
15.A New Species and the First Record of the Genus Anillinus (Carabidae: Trechinae: Bembidiini) from the Ozark Region
Igor M. Sokolov and Larry E. Watrous. 2008. Coleopterists Bulletin.
16.American Beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia
Arnett, R.H., Jr., and M. C. Thomas. (eds.). 2000. CRC Press LLC, Boca Raton, FL.
17.Evolution of the (Coleoptera) Carabidae of the Southern Appalachians.
Thomas C. Barr Jr. . 1969. Virginia Polytechnic Institute .