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Genus Anomala

Scarab - Anomala digressa Shining Leaf Chafer Beetle - Anomala Shining Leaf Chafer - Anomala marginata BG2386 D2887 - Anomala undulata beetle id - Anomala marginata Scarab - Anomala Which small brown beetle? - Anomala Maybe a Shining Leaf Chafer - Anomala
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Scarabaeidae (Scarab Beetles)
Subfamily Rutelinae (Shining Leaf Chafers)
Tribe Anomalini
Genus Anomala
Synonyms and other taxonomic changes
selected synonyms:
Phyllopertha Stephens 1830
Callistethus Blanchard 1851 [used for marginata by Ratcliffe & Paulsen (1)]
Pachystethus Blanchard 1851 [was used in much of the older NA literature for A. marginata, lucicola & oblivia]
Anomala (Paranomala) Casey, 1915
Paranomala Casey, 1915 (2) - widely used in GBIF and iNat, but mostly for spp. occurring in Mexico to S. America (3)
Exomala Reitter 1903 [often seen used for A. orientalis]

revised in Ramírez-Ponce and Morón, 2009 (2)
Explanation of Names
Anomala Samouelle 1819
Latin anomala 'irregular, deviant'
48 spp. in our area(4), 180 in the New World; a huge group, with >1,000 described species worldwide (some of questionable validity)(5)
6-12 mm
Some characteristics of the genus:
small to medium-sized, slightly oval-shaped, convex in cross-section
antennae with 9 segments
pronotum not pubescent (hairy)
elytra with shallow striations and punctures, rather translucent (membranous) along sides
Species identification
Mesosternum between mesocoxae concave to flat anteriorly, not elevated posteriorly to more than low umbo ......... top, most spp.
Mesocoxae separated by distinctly convex to prominently raised mesosternal process, sometimes porrect ...... bottom, includes spp. such as A. marginata, lucicola & oblivia [These were placed in the genus Pachystethus in much of the older American literature and Callistethus was used in combination with marginata in(1)]

Key to NA species in(6)
Anomala (sensu lato) Samouelle 1819 - world wide distribution - Map (3)
Paranomala Casey, 1915 - e. US to s. Calif. / Mex. to Columbia - Map (3)
Casey (1915) divided the genus Anomala into 3 groups with subgenus status. He called the first Paranomala and included most of the species studied in it. That name was synonymized by Machatschke (1957) with Anomala. (2)
Print References
Casey, T.L. 1915. Memoirs on the Coleoptera. VI. New Era Printing Company, Lancaster, PA. 460 pp.
Machatschke, J. 1957. Coleoptera Lamellicornia, Scarabaeidae, Rutelinae, Anomalini. Genera Insectorum, fasc. 199B: 1–219.
Ramírez-Ponce, A. and M.A. Morón. 2009. Relaciones filogenéticas del género Anomala Samouelle (Coleoptera: Melolonthidae: Rutelinae: Anomalini). Rev. Mex. Biodivers. 80: 357–394.
Samouelle, G. 1819. The entomologist's useful compendium or an introduction to the knowledge of British Insects. Thomas Boys, London. 496 pp.
Works Cited
1.The Scarabaeoid Beetles of Nebraska
Brett C. Ratcliffe & M.J. Paulsen. 2008. University of Nebraska State Museum, Vol 22, 570 pp.
2.Relaciones filogenéticas del género Anomala Samouelle (Coleoptera: Melolonthidae: Rutelinae: Anomalini).
Ramírez-Ponce, A. and M.A. Morón. 2009. Revista Mexicana de Biodiversidad 80(2): 357–394.
3.Global Biodiversity Information Facility (GBIF)
4.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
5.Generic Guide to New World Scarab Beetles (by Brett Ratcliffe and Mary Liz Jameson)
6.Revision of the Scarabaeidae: Anomalinae. 3. A key to the species of Anomala of America north of Mexico
R.W. Potts. 1977. The Pan-Pacific Entomologist, 53: 129-134.
7.Scarab beetles (Coleoptera: Scarabaeidae) of South Carolina
Phillip J. Harpootlian. 2001. Clemson University Public Service.