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Entypus
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Entypus unifasciatus
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Entypus unifasciatus californicus
Photo#528147
Copyright © 2011
Robyn Waayers
- -
Entypus unifasciatus
-
Lake Sutherland, San Diego County, California, USA
June 18, 2004
Size: 19 mm
Found on stinging nettle (Urtica dioica) near a reservoir in the foothills. Chamise chaparral and oak woodlands surroundings. 627 m.
Images of this individual:
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Contributed by
Robyn Waayers
on 12 June, 2011 - 5:38pm
Last updated 16 October, 2015 - 1:40pm
Moved
Moved from
Pepsini
.
…
Robyn Waayers
, 16 October, 2015 - 1:11pm
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Another genus (other than Hemipepsis or Pepsis)?
Robyn...looking at your full-size image...note that the 1st recurrent vein meets the 2nd submarginal cell at
about 1/4
its length away from its distal end. (It's often tricky to "see" this without careful scrutiny...because the shallow angle at which the veins meet gives the impression of them being "parallel near their intersection"...and the human eye/mind seems to like to (inaccurately) interpret that as if the meeting point were closer than it really is to the junction of the two adjacent veins bounding the postero-proximal corner of the 3rd submarginal cell!)
The above suggests this isn't
Hemipepsis
(where the 1st recurrent vein should meet within
a tenth
of the length of the 2nd submarginal cell from its distal end).
I haven't tried to run this through the keys (to genera and species) in Townes
(
1
)
(1957) yet, but I'm wondering if it may be
Entypus
? If not, I guess it would be one of the other (yet smaller) pepsine genera. It's good to have a specimen at hand in trying to work through those keys, as they often lists characters that aren't be discernible in photos.
…
Aaron Schusteff
, 7 October, 2015 - 12:51am
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Almost forgot that I had posted this image!
I was thinking of trying to work on these guys (I have three like them) after starting to feel more confident about Pepsis and Hemipepsis, and I think you're right - they might be from another genus altogether. Will revisit when the school week settles down. I have this specimen so that might help, as you say!
…
Robyn Waayers
, 7 October, 2015 - 10:37am
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"Almost forgot"
Good point...I hadn't noticed how long ago it was since you'd found that wasp! Nice that you still have the specimen. And not only that, but nice it was prepared so well (i.e. spread, etc.). Working with a reference specimen like that will be a pleasure!
(Then again,
all
your specimens are beautifully prepared! :-)
…
Aaron Schusteff
, 7 October, 2015 - 4:15pm
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Thanks : )
This does look like Entypus based on the venation pattern you describe elsewhere on the guide (based on where 1m-cu meets S2 - around 1/4 - or in this case, more like 1/3 - of the way from distal end). E. unifasciatus californicus seems to be the only one on BG with orange antennae and entirely orange wings. I re-uploaded a slightly lightened photo, too. I'm adding a close-up of the wing venation, too.
…
Robyn Waayers
, 9 October, 2015 - 3:33pm
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I tried working from your photos
...using Townes
(
1
)
and the
online key to genera of New World Pepsinae
from the "Pompilid Project".
And I pretty much got the same result,
E. unifasciatus californicus
. I left a comment under the post below for Nick Fensler, to hopefully get his ideas/insights here:
…
Aaron Schusteff
, 13 October, 2015 - 9:20am
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.
Thanks Aaron!
…
Robyn Waayers
, 13 October, 2015 - 10:52am
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More Details
In working through the online
Key to the New World Genera of Pepsinae
at the "Pompilid Project" web site, and the key to the genera of Pepsinae in Townes
(
1
)
, I was able to use info visible in the two photos here to narrow the candidate genera down to
Chirodamus (=Calopompilus), Priocnessus
, or
Priocnemioides (= Entypus)
. Note that the two "Townes names"
Chirodamus
and
Priocnemioides
have current synonyms
Calopompilus
and
Entypus
, respectively. The later names are those used on BugGuide and in the key at the "Pompilid Project" web site. But I'll be referring to genus and species descriptions and range maps only found in Townes
(
1
)
below, and so will often use his names here to make it easier to cross-reference with his monograph.
Now, from reading Townes' genus description for
Chirodamus
, I do
not
think that's what we've got here...due to wing venation discrepancies. But to positively eliminate that genus using Townes' key, it would suffice to get good views of:
The clypeus...showing whether it's "wide, short, and rather flat" (
Chirodamus
) vs. "longer and more convex" (
Priocnessus
or
Priocnemioides
), and;
The "brush" on inner side of hind tibia...showing whether it's "broadly continuous to the apex" (
Chirodamus
) vs. "with a sub-apical constriction or interruption" (
Priocnessus
or
Priocnemioides
)
Assuming this is not
Chirodamus
, then at couplet 5 of Townes key, details of wing venation clearly visible in the photos here (i.e. "Cubital vein not quite reaching the wing margin and the nervellus ending at or distad of the juncture of cubitella with discoidella") take this to
Priocnessus
rather than
Priocnemioides
.
But when I take your photo through the key to species for
Priocnessus
, then the clearly evident characters "Head and body black" (vs. "rufous or mostly rufous") and "Wings orange" (vs. "subhyaline to blackish") leads me promptly to
Priocnessus nigricans
...which is given as a species from southeastern Arizona. Moreover, the 1st recurrent vein in the photo meets the 2nd submarginal cell near its distal 1/4, whereas Townes
(
1
)
states the meeting point is "
at or just
beyond its middle" in his genus description for
Priocnessus
. The genus description also states that
Priocnessus
is "of rather slender build", whereas I'd say the body here is somewhat stocky. So I suspect the key to genus may have led us astray, and that this may not be
Priocnessus
.
The final alternative for genus using Townes
(
1
)
is what he called
Priocnemioides
, which is now referred to as
Entypus
. Indeed Townes genus description for
Priocnemioides
fits the wasp here very well. As far as the the species key goes, the characters in the first couplet are not visible in the photos here (we'd need a view of the "brushes" on the inner hind tibia; and/or the "nipples" on posterior part of apical margin of fore coxa; and/or the second sternite of female). But if one follows the first choice in couplet 1 and proceeds to couplet 2...there is an immediate obstruction to going any further along that route, namely:
2a. Flagellum orange; wings black......
magnus
2b. Flagellum black; wings orange to black......3
Neither 2a nor 2b hold here...so it seems safe to assume we should follow the second lead of couplet 1...and proceed to couplet 7. Doing so leads without complications to
E. unifasciatus californicus
, which besides agreeing well with all the key characters, is also the only species of
Priocnemioides (= Entypus)
shown as occurring in California in Townes range maps...except
E. texanus texanus
, which has black antennae.
Since the reasoning above leading to
Entypus
depended on eliminating alternatives for genera and species, it could be rendered incorrect if other species of
Entypus
or
Priocnessus
have been described since Townes 1957 monograph. But
this current checklist
from the "Pompilid Project" web site lists exactly the same taxa for
Entypus
that Townes treated under the genus name of
Priocnemioides
. So if this is indeed
Entypus
, it would seem to have to be
E. unifasciatus californicus
. As for
Priocnessus
, the species on the checklist include all those treated in Townes
(
1
)
, together with exactly one additional species: namely,
Priocnessus hurdi
Dreisbach, 1960. A key including
P. hurdi
appears
here
, and there is at least
one specimen at the Essig Museum
. If need be, eventually I can try to look at the specimen next time I go to the Essig Museum (or maybe we can persuade Joyce to do so, as she works around the corner from the museum :-).
So at this point, as far as I can tell, this is either
E. unifasciatus californicus
or
P. hurdi
. And I think we'll eventually be able to settle the ID here.
…
Aaron Schusteff
, 14 October, 2015 - 4:33am
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No...
this is
Entypus
or
Cryptocheilus
. Those two genera are very similar and possibly should not be separated (in which case all species of
Entypus
would be transferred to
Cryptocheilus
). It is definitely not
Priocnessus
or
Calopompilus
. There are two species of
Cryptocheilus
with orange wings and flagellae:
C. attenuatum
and
C. pallidipenne
. Please note that the geographic ranges in Townes are almost meaningless (especially for
C. attenuatum
).
…
Nick Fensler
, 14 October, 2015 - 12:07pm
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Good to get your feedback, Nick
And I'm thinking your "No" is also a partial "Yes"...in that you, Robyn, and I all seem to be circling around, and gravitating towards,
Entypus
...but with different degrees and directions of uncertainty.
Perhaps it wasn't clear, but my concluding thoughts in my previous comment were that this is
most likely
Entypus
(and if so,
E. unifasciatus californicus
)...but (at that time) I could not absolutely rule out the (unlikely) possibility of
Priocnessus hurdi
(for the reasons given in the 3rd and next-to-last paragraph of that comment).
However, today I was able to study the original description of
P. hurdi
in
Dreisbach(1960)
, and I can now confidently rule out that species here..as the female is described as having body largely rufous. So we're now both on the same page as far as this
not
being
Priocnessus
or
Calopompilus
(me mostly from "book knowledge", and you from that plus more time/experience studying specimens).
So now, how about the question of
Entypus
vs.
Cryptocheilus
here? Both Townes
(
1
)
(couplet 7 on pg. 11) and the
"Pompilid Project" key
(at couplet 9 there) use the same pair of characters to separate these two groupings, Townes' version being (see Fig. 1A on pg. 7 for wing venation terminology):
5a)
Carina on
mesosternum
in front of each middle coxa angled medially and, at the angle, usually produced as a tooth
; nervellus
ending beyond, at, or just before the juncture of cubitella with discoidella ......
Entypus
5b)
Carina on
mesosternum
in front of each middle coxa evenly curved
; nervellus
ending distinctly before the juncture of cubitella with discoidella.....
Cryptocheilus
While assessing the primary "mesosternum character" in the couplet above would require a well-lit and resolved close-up image of the metasternum (and those in the
"Pompilid Project" key
for couplet 9 are not as clear as I'd like), I think the secondary "hind wing venation" character...involving the meeting point of the nervellus with the cubitella and discoidella...clearly indicates
Entypus
here.
Assuming this is
Entypus
, to get through the 1st couplet in the key to species (on pg. 50 of Townes
(
1
)
), a good underside view (as you commented
here
) would help decide which of the two leads to go with. But if you look ahead to couplet 2 there, you can see that neither lead works for the female here...so it seems (indirectly) clear that one must go with couplet 7. And from there, this keys unambiguously to
E. unifasciatus californicus
.
If this were
Cryptocheilus
, then (as you mentioned above) the key to species (on pg. 69 of Townes
(
1
)
) indicates either
C. attenuatum
or
C. pallidipenne
, due to the orange antennae here. And the ranges given for those two species ("Tennessee, Louisiana, Texas, and Iowa" for the former, versus "Arizona and California" for the latter) are quite disjunct and would strongly suggest
C. pallidipenne
. Even taking into account your skepticism of the dependability of the range maps in Townes...this still gives at least a suggestion of higher likelihood for
pallidipenne
way out west in CA.
Similarly, if you scrutinize Townes' range map for
E. unifasciatus californicus
(on pg. 65), you'll see a large cluster of collection record dots in the vicinity of Robyn's coastal mountain location here...whereas the California records shown on the range map for
C. pallidipenne
(on pg. 74) are all clustered in the desert areas of far southeast CA.
Overall, I'd say things point to
E. unifasciatus californicus
here.
…
Aaron Schusteff
, 14 October, 2015 - 11:55pm
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Have had a chance to look over...
...and think about your analysis, Aaron. I see the hindwing character. I decided to move it. Many thanks!
…
Robyn Waayers
, 16 October, 2015 - 1:03pm
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