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Tribe Athysanini

Spotted hopper - Colladonus geminatus Leafhopper - Allygus mixtus Cicadellid - Deltocephalinae? - Thamnotettix zelleri Unknown Leafhopper - Colladonus montanus Yellow leafhopper - Colladonus geminatus orange/brown hopper at light - Norvellina helenae Leafhopper - Mesamia Eutettix - Eutettix nitens
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Auchenorrhyncha (True Hoppers)
Infraorder Cicadomorpha (Cicadas, Spittlebugs, Leafhoppers, and Treehoppers)
Superfamily Membracoidea (Leafhoppers and Treehoppers)
Family Cicadellidae (Typical Leafhoppers)
Subfamily Deltocephalinae
Tribe Athysanini
Synonyms and other taxonomic changes
Phrynomorphini Kirkaldy 1907, Thamnotettigini Distant 1908, Euscelini Van Duzee 1917, Anoterostemmini Haupt 1929, Bobacellini Kusnezov 1929, Platymetopiini Haupt 1929, Colladonini Bliven 1955, Cerrillini Linnavuori 1975, Allygidiina Dmitriev 2006
42(?) genera in our area, >1800 spp. in ~300 genera total(1)
Informal clades based on genetic analyses:
Athysanus clade
Athysanus Burmeister 1838
Streptanus Ribaut 1942
Euscelis clade
Allygidius Ribaut 1948
Allygus Fieber 1872
Euscelidius Ribaut 1942
Euscelis Brullé 1832
Extrusanus Oman 1949
Graphocraerus Thomson 1869
Thamnotettix Zetterstedt 1838
untested—Athysanus or Euscelis clade
Conosanus Osborn & Ball 1902
Coulinus Beirne 1954
Doliotettix Ribaut 1942
Ederranus Ribaut 1942
Macustus Ribaut 1942
Colladonus clade
Bonneyana Oman 1949
Colladonus Ball 1936
Idiodonus Ball 1936
Brazosa clade
Zabrosa Oman 1949
Norvellina clade
Aplanus Oman 1949
Bandara Ball 1931
Eusama Oman 1949
Fitchana Oman 1949
Mesamia Ball 1907
Norvellina Ball 1931
Atanus clade
Atanus Oman 1938
Eutettix clade
Aligia Ball 1907
Crassana DeLong & Hershberger 1947
Eutettix Van Duzee 1892
Ollarianus Ball 1936
Orientus clade
Orientus DeLong 1938
Aplanusiella Nielson & Haws 1992
Caladonus Oman 1949
Cetexa Oman 1949
Danbara Oman 1949
Eusora Oman 1949
Floridonus Oman 1949
Fridonus Oman 1949
Hesium Ribaut 1942
Nigridonus Oman 1949
Nurenus Oman 1949
Sobara Oman 1949
Twiningia Ball 1931
nearly all terrestrial ecosystems
wide variety of eudicots and occasionally grasses and sedges
The largest, highly polyphyletic tribe of Deltocephalinae, pretty much the "Incertae sedis" Deltocephalinae. It was previously broken up into a number of subtribes which were genera grouped based on morphology—DNA and phylogenic analysis has revealed that none of those subtribes are valid except for Cicadulina which was elevated to tribal status after the group was proved to be monophyletic. The Athysanus-group is an informal genus group that is mostly monophyletic and groups all species related to the genus Athysanus. This genus group is likely the eventual "true" Athysanini after the tribe is revised. Some members of Athysanini may eventually belong to existing tribes, but most may end up in new tribes. The other large tribes (Deltocephalini, Paralimnini) are generally understood to be monophyletic.