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Genus Nallachius

Pleasing Lacewing - Nallachius americanus - male Pleasing Lacewing - Nallachius americanus - male Pleasing Lacewing? - Nallachius americanus - male Unfamiliar bug - Nallachius americanus - male Hexapoda - Nallachius americanus - male 20170611-DSC_4255 - Nallachius americanus - female Dilaridae - Nallachius americanus unknown bug - Nallachius americanus
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Neuroptera (Antlions, Owlflies, Lacewings, Mantidflies and Allies)
Suborder Hemerobiiformia (Lacewings, Mantidflies and Allies)
Family Dilaridae (Pleasing Lacewings)
Genus Nallachius
Explanation of Names
Nallachius Navás 1909
Numbers
2 spp. in our area(1):
  N. americanus
  N. pulchellus
Identification
Larvae very elongate:
     
Adults resemble small brown moths: wings widely-rounded, hairy; costal cross veins of wings not forked; male antennae pectinate, female filiform; female ovipositor long, recurved over abdomen:
      male N. americanus       female N. pulchellus
Our two species can be separated by details of wing venation; genitalia; and range...see keys and discussions in Penny(2002); Adams(1970); or Carpenter(2)(1940). Penny(3)(2002) gives detailed wing venation diagrams for N. americanus and N. pulchellus.
    Another wing venation diagram for N. americanus below:
       
Range
  N. americanus: e. US (NJ-FL to MI-KS-TX),
  N. pulchellus: AZ(1)(4)(BG data)
Habitat
larvae under tree bark; adults come to light
Food
Larvae are presumed to feed on larvae & eggs of various tree-boring or bark-infesting beetles and ants.
Remarks
Uncommon. Adults are most often collected on or in close proximity to dead or living trees: in particular tulip poplar (Liriodendron tulipifera); and several species of oaks (Quercus).
Print References
Adams, P. A. (1970). A review of the New World Dilaridae. Postilla 148:1-30. (Full Text)
Bowles, D.E., A. Contreras-Ramos, M. A. Sarmiento-Cordero, & M. L. Ferro (2015). New distributional records for pleasing lacewings (Neuroptera: Dilaridae, Nallachius spp.) in the Americas. Insecta Mundi 0406: 1–10 (Full Text)
Carpenter, F. M. (1940). A revision of the Nearctic Hemerobiidae, Berothidae, Sisyridae, Polystoechotidae and Dilaridae (Neuroptera). Proc. Amer. Acad. Arts and Sci. 74:193-260.(2)
Carpenter, F.M. (1947). Taxonomic Notes on the Dilaridae (Neuroptera). Psyche 54: 100-109 (Full Text)
Gurney, A. B. (1947) Notes on Dilaridae and Berothidae, with special reference to the immature stages of the Nearctic genera (Neuroptera). Psyche 54:145-169 (Full text)
Hoffman K.M. (1990) Distributional notes on North and Central American Dilaridae. Entomological News 101: 155-157. (Full Text)
Kuhar, T. P. (1995). A pleasing lacewing, Nallachius americanus (McLachan), from southwestern Virginia (Neuroptera: Dilaridae). Banister 5: 32–33. (Full Text)
MacLeod E.G., Spiegler P.E. (1961) Notes on the larval habitat and developmental peculiarities of Nallachius americanus (McLachlan) (Neuroptera: Dilaridae). Proc. Ent. Soc. Wash. 63: 281-286. (Full Text)
Monserrat V.J. (2005) Nuevos datos sobre algunas pequeñas familias de neurópteros (Insecta: Neuroptera: Nevrorthidae,. Osmylidae, Sisyridae, Dilaridae). Heteropterus Revista Entomologia 5: 1–26. (Full text, see pg. 18 for Nallachius)
Oswald, J. D., (1998). Annotated Catalogue of the Dilaridae (Insecta: Neuroptera) of the World. – Tijdschrift voor Entomologie 141: 115-128. [ISSN 0040-7496] (PDF).
Preny, T. J. & R. J. L. Jones (2017). A new family to Canada with the discovery of the Pleasing Lacewing Nallachius americanus (McLachlan) (Neuroptera: Dilaridae) at the Ojibway Prairie Complex in Windsor, Ontario. J. ent. Soc. Ont. 148: 39–41. (Full Text)
Sperka, C. L. (1972). Nallachius americanus (McLachlan) (Neuroptera: Dilaridae) in Kentucky. Entomological News 83: 282–283. (Full Text)
Stange, L. (1961). New records for the genus Nallachius Navas. Pan-Pacific Entomologist 38(2):144. (Full Text)
Steyskal, G.C. (1944) Notes on Nallichius americanus. Psyche 51:183-184 (Full text)
Internet References
Katja Schulz on Flickr -- Remarkable oviposition videos: #1 and #2; two mating photos #1, #2; 6 oviposition photos (scroll w/ left arrow through photostream); 4 photos (of pheromone broadcasting?); and 4 photos of mating pair.
Species page for N. americanus from North Carolina State University Insect Museum
Illinois record for N. americanus...see "A Rare Jewel" (from The Illinois Natural History Survey reports, No. 60, October 1967).
Virginia record for N. americanus in 1995 article by T.P. Kuhar (of Virginia Polytechnic Institute and State U.)
Works Cited
1.Species catalog of the Neuroptera, Megaloptera, and Raphidioptera of America North of Mexico
Penny N.D., Adams P.A., Stange L.A. 1997. Proc. Calif. Acad. Sci. 50: 39-114.
2.A revision of the Nearctic Hemerobiidae, Berothidae, Sisyridae, Polystoechotidae and Dilaridae (Neuroptera)
Carpenter F.M. 1940. Proc. Amer. Acad. Arts and Sciences 74: 193-280.
3.A guide to the lacewings (Neuroptera) of Costa Rica
Norman D. Penny (author & editor), Kevin M. Hoffman, Martin Meinander, Victor J. Monserrat, Lionel A. Stange. 2002. Proceedings of the California Academy of Sciences 53(12): 161-457.
4.A Checklist and Bibliography of the Megaloptera and Neuroptera of Florida (by L. Stange)