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Family Geotrupidae - Earth-Boring Scarab Beetles

BG2096 C1997 - Bolbocerosoma farctum Bolbocerosoma? - Bolbocerosoma bruneri Bolbo - Bolborhombus sallei Eucanthus lazarus (Fabricius) - Eucanthus lazarus Scarab beetle - Geotrupes opacus Geotrupidae? - Odonteus Geotrupes Beetles - Geotrupes splendidus
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Geotrupidae (Earth-Boring Scarab Beetles)
Synonyms and other taxonomic changes
Until recently treated as a subfamily of Scarabaeidae
Explanation of Names
Geotrupidae Latreille 1802
Numbers
55 spp. in 11 genera in the Nearctic(1), ~620 spp. in ~70 genera worldwide(2) arranged into 13 tribes and 3 subfamilies(3)
Overview of our fauna:
Family GEOTRUPIDAE
Subfamily BOLBOCERATINAE
Odonteus 10 spp., mostly eastern
Eucanthus 5 spp., mostly eastern
Bradycinetulus 3 spp., eastern/central US
Bolbocerastes 3 spp., w. US
Bolbocerosoma 10 spp., eastern/central US
Bolbelasmus 4 spp., sw. US
Bolborhombus 3 spp., sw. US
Subfamily GEOTRUPINAE
Geotrupes 9 spp., US except far west
Mycotrupes 5 spp., se. US
Peltotrupes 2 spp., Florida
Ceratophyus 1 sp., California
Size
5-45 mm(2)
Identification
Body stout, often hemispherical; antenna 11-segmented (Scarabaediae have fewer segments)

keys in(4)
Subfamilies:
Geotrupinae (left): Antennal club small, body largely black with or without metallic sheen
Bolboceratinae (right): Antennal club large, body brown, some with black markings
Habitat
These beetles spend most of their lives in burrows one to four feet down, often under dung or carrion.
Food
Larvae feed on dung or carrion; adults, on dung or fungi, or not at all
Internet References
Works Cited
1.American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea
Arnett, R.H., Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). 2002. CRC Press LLC, Boca Raton, FL.
2.Generic Guide to New World Scarab Beetles (by Brett Ratcliffe and Mary Liz Jameson)
3.Family-group names in Coleoptera (Insecta)
Bouchard P. et al. 2011. ZooKeys 88: 1–972.
4.The Geotrupinae of North and Central America
Howden H.F. 1964. Mem. Ent. Soc. Canada 39: 1-91.