Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Family Geotrupidae - Earth-Boring Scarab Beetles


The Geotrupinae of North and Central America
By Howden H.F.
Mem. Ent. Soc. Canada 39: 1-91, 1964

Biology and taxonomy of North American beetles of the subfamily Geotrupinae, with revisions of the genera....
By H.F. Howden
Proceedings of the United States National Museum, 104(3342): 151-319, plates 1-18., 1955
Full title: "Biology and taxonomy of North American beetles of the subfamily Geotrupinae, with revisions of the genera Bolbocerosoma, Eucanthus, Geotrupes, and Peltotrupes (Scarabaeidae)"

Online here.

Revision of the genus Odontaeus, Dej.
By Wallis J.B.
Can. Entomol. 60: 119‒128; 151‒156; 168‒176, 1928

An illustrated key to the genus Geotrupes Latreille (Coleoptera: Geotrupidae) of Maryland including notes on G. ulkei...
By Guarnieri F.G., Harpootlian P.J.
The Maryland Entomologist 6(1): 6–13, 2013
Full title: An illustrated key to the genus Geotrupes Latreille (Coleoptera: Geotrupidae) of Maryland including notes on G. ulkei Blanchard, a rare southern species recently discovered in Morgan County, West Virginia

very helpful paper; i have a copy, courtesy Phil Harpootlian (both authors are long-time BG contributors)

Scarabaeid beetles of the genus Bradycinetulus and closely related genera in the United States
By O. L. Cartwright
Proceedings of the United States National Museum, Washington Vol. 103, No. 3318: 95-120, 1953
O. L. Cartwright, Scarabaeid beetles of the genus Bradycinetulus and closely related genera in the United States, Proceedings of the United States National Museum, Washington 103(3318):95-120, 1953.

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Aquatic Hydrophilidae and Hydraenidae of Wisconsin (Coleoptera). I.
By William L. Hilsenhoff
The Great Lakes Entomologist, 1995
Full title: Aquatic Hydrophilidae and Hydraenidae of Wisconsin (Coleoptera). I. Introduction, Key to Genera of Adults, and Distribution, Habitat, Life Cycle, and Identification of Species of Helophorus Fabricius, Hydrochus Leach, and Berosus Leach (Hydrophilidae), and Hydraenidae

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Saprophagous Scarabaeidae (Coleoptera) of North Dakota
By Helgesen R.G., Post R.L.
Schafer-Post Series: North Dakota Insects Publication 7, NDSU. 60 pp., 1967

The phytophagous Scarabaeidae and Troginae (Coleoptera) of North Dakota
By Lago P.K., Post R.L., Oseto C.Y.
Schafer-Post Series: North Dakota Insects Publication 12, NDSU. 131 pp., 1979