Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Subtribe Bembidiina


Phylogeny of Bembidion and related ground beetles (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidiina)
By Maddison D.R.
Mol. Phylogenet. Evol. 63: 533‒576, 2012

Rediscovery and distribution of Bembidion plagiatum Zimmerman (Coleoptera: Carabidae)
By Richard L. Hoffman
Brimleyana No.7 pp145-150, 1982
Online Here.

Detailed discussion of B. plagiatum's habitat preference and distribution. Prior to the publication of this note, the species was very poorly known and rare in collections.

Re-collected after 55 years: a new species of Bembidion (Coleoptera, Carabidae) from California
By Maddison D.R., Sproul J.S., Will K.
ZooKeys 1156: 87–106, 2023

Bembidion ambiguum (Coleoptera: Carabidae) is established in California
By D.R. Maddison, K. Will, S. Crews, J. LaBonte
Biodiversity Data Journal 6: e30763, 2018
First records of Bembidion ambiguum in North America presented, along with diagnostic photographs and modified key couplets.
Available online as of November 2020.

Systematics of the north american beetle subgenus Pseudoperyphus (Coleoptera: Carabidae: Bembidion) [...]
By David R. Maddison
Annals of Carnegie Museum, Vol 77 No 1 pp.147-193, 2008
Full title: "Systematics of the north american beetle subgenus Pseudoperyphus (Coleoptera: Carabidae: Bembidion) based upon morphological, chromosomal, and molecular data"

A masterful treatment of this immensely difficult group of Bembidion species, with desciption of four new species, detailed illustrations of external morphological characters as well as characters of the aedeagus, discussion on range and habitat preference, and notes on future problems that remain to be solved.

Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Col.: Carabidae)
By Maddison D.R., Sproul J.S.
Zool. J. Linn. Soc. XX: 1-46, 2020

Shards, sequences, and shorelines: two new species of Bembidion from North America (Coleoptera, Carabidae)
By David R. Maddison
ZooKeys 1007: 85–128, 2020
Good descriptions of some species and additions / corrections to Lindroth's keys.

PDF here.

A preliminary characterization of Bembidion perspicuum LeConte, with a reclassification of related species (Coleoptera, Carabida
By David R. Maddison, Alexander P. Swanson
ZooKeys 43: 15-31, 2010
Good descriptions of some species and additions / corrections to Lindroth's keys.

PDF here.