Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada

Genus Harpalus


The taxonomy and speciation of Pseudophonus
By Ball, G.E. and J.N. Anderson
The Catholic University of America Press, Washington, D.C. XI + 94 pp., 1962
Limited online access here.

Classification of the species of the Harpalus subgenus Glanodes Casey (Carabidae: Coleoptera)
By Ball G.E.
Col. Bull. 26: 179–204, 1972

Review of the species of the subgenus Megapangus
By Will K.W.
Col. Bull. 51: 43‒51, 1997

Classification of the genus Harpalus (Coleoptera, Carabidae) of the World based on imaginal morphology
By Kataev B.M.
Diversity 15, 971. 137 pp., 2023

A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new ...
By Danny Shpeley, Wesley Micheal Hunting, George E. Ball
ZooKeys 690: 1-195 , 2017
Full title: A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography
Full text(PDF)

An annotated checklist of ground beetles (Carabidae) in the state of Sonora, Mexico with new species & collection localities
By Spheley, D & T.R. Van Devender
Dugesiana 29(1): 41-137, 2021
Complete title: An annotated checklist of ground beetles (Insecta: Coleoptera: Carabidae) occurring in the state of Sonora, Mexico with descriptions of new species and notes about some collection localities.

ABSTRACT
An annotated checklist of ground beetles in the family Carabidae from the state of Sonora in Mexico is presented with notes including number of specimens examined, municipios, habitat, vegetation, elevation, and collection dates. The checklist includes 388 taxa, of which 270 are described taxa (included are 19 subspecies and 1 hybrid) and 118 are unnamed morphospecies, in 92 genera representing 35 tribes. Dot maps display where specimens were collected. Three new species are described, with corrected keys and images: Calathus mcclevei, new species (type locality: Mexico, Sonora, Municipality of Yécora, 10 km south of Yécora, ca. 28.278° N 108.926° W); Calathus tigrinus, new species (type locality: Mexico, Sonora, Municipality of Nacozari de García, Rancho El Tigre, 28.3 km (by air) WNW Bavispe, Sierra El Tigre, 30.58988° N 109.20811° W); and Amblygnathus balli, new species (type locality: Mexico, Sonora, Municipality of Fronteras, Rancho Capulín, 23.5 km (by air) WSW Fronteras, Sierra Buenos Aires, 30.76167° N 109.82361° W).

Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi (new species), Zuphioides (new genus), and classification
By Ball, G.E. and D. Shpeley
ZooKeys 315: 17-54 , 2013
Full Title: Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi, new species, and Zuphioides, new genus, and classification of the genera (Coleoptera, Carabidae)

Downloadable at DOI: 10.3897/zookeys.315.5293

The article's main impact on BugGuide is that North American species previously belonging to Zuphium would be moved to genus Zuphioides. The latter Latinized name is treated as neuter and so the previous epithets under Zuphium remain unaffected.

Species delimitation in the ground beetle subgenus Liocosmius (Bembidion), including standard and next-generation sequencing
By Maddison, DR & KW Cooper
Zoological Journal of the Linnean Society, 2014
Full title: Species delimitation in the ground beetle subgenus Liocosmius (Coleoptera: Carabidae: Bembidion),
including standard and next-generation sequencing of museum specimens

DAVID R. MADDISON & KENNETH W. COOPER

Zoological Journal of the Linnean Society, 2014, 172, 741–770. With 19 figures