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Species Harpalus actiosus

Harpalus actiosus Casey - Harpalus actiosus - female
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Harpalinae
Supertribe Harpalitae
Tribe Harpalini
Subtribe Harpalina
Genus Harpalus
No Taxon (Subgenus Pseudoophonus)
No Taxon (compar group)
Species actiosus (Harpalus actiosus)
Explanation of Names
Harpalus actiosus Casey 1914
16 mm

Det. P. W. Messer, 2013 (by microscopic examination)
Characteristic combination:
Elytra densely & uniformly punctate
anterior pronotal border slightly angularly emarginate (difficult to see in photo)
large size (ABL ~ 16 mm)
Texas location.
This combination rules out the two most similar species: H. protractus and H. vagans (formerly H. longicollis).
NJ-VA-MS to ND-CO-TX (1)
Print References
Casey, T.L. 1914. Memoirs on the Coleoptera. V. New Era Printing Company, Lancaster. 387 pp.
Works Cited
1.Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico
Bousquet Y. 2012. ZooKeys 245: 1–1722.