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Species Harpalus paratus

Harpalus paratus Casey - Harpalus paratus - female
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Adephaga (Ground and Water Beetles)
Family Carabidae (Ground Beetles)
Subfamily Harpalinae
Supertribe Harpalitae
Tribe Harpalini
Subtribe Harpalina
Genus Harpalus
No Taxon (Subgenus Pseudoophonus)
No Taxon (compar group)
Species paratus (Harpalus paratus)
Explanation of Names
Harpalus paratus Casey, 1924
ABL ~ 16 mm

Det. P. W. Messer, 2013 (based on microscopic examination of this specimen)
Distinguished from juxatapostional species in Ball & Anderson (1962) by combination of:
large size (elytra length = or > 8 mm)
elytra without (or rare) punctulae
hind tarsi dorsally impunctate and glabrous (or rare setae).
The latter diagnostic character I find especially useful. Other large Pseudoophonus have hind tarsi with obvious dorsal pubescence or at least show numerous punctulae under high power. (per Peter W. Messer, 26 January, 2013)
AZ-TX-OH-ID (+PA?) / AB-SK / Mex. (1)
primarily a prairie species (1)
Print References
Ball, G.E., and J.N. Anderson. 1962. The taxonomy and speciation of Pseudophonus (a subgenus of Harpalus: Harpalini: Carabidae, known to occur in North America). Studies on Speciation No. 1. The Catholic University of America Press, Washington D.C. xi + 94 pp.
Casey, T.L. 1924. Memoirs on the Coleoptera. XI. New Era Printing Company, Lancaster. 347 pp.
Works Cited
1.Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico
Bousquet Y. 2012. ZooKeys 245: 1–1722.