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Registry/Forum for Tracking New Records of Nearctic Ground Beetles (Geadephaga)

Keywords: Caraboidea, Carabidae, carabid, Tiger Beetles, new species, taxonomy, North America, United States, Canada

The large beetle group designated "Geadephaga", commonly known as the "ground beetles", is the superfamily Caraboidea composed of the three families Trachypachidae, Rhysodidae, and Carabidae. The latter family is by far the largest one. Its members, informally named "carabids", account for 99.55 percent of the roughly 2700 known species in the Nearctic Caraboidea north of Mexico. It is this faunal region that concerns BugGuide and our post-2012 Caraboidea Registry/Forum. The family Carabidae includes several subfamilies of which Cicindelinae is represented by the popular "Tiger Beetles". See the comment below regarding the state of Tiger Beetle taxonomy.

This frequently-updated website is an interactive clearinghouse where users of provide new information to supplement the geographic distributions and taxonomies published in the massive 2012 catalogue of Nearctic Geadephaga by Yves Bousquet (1) [YB2012]. Newly documented states or provinces are entered by their standard abbreviations after the alphabetically-listed species names below. In addition, geographic records that confirm the questionable ones enclosed in square brackets by Bousquet, will be annotated here as verified. In tracking the taxonomic literature, we will include Nearctic species that are new to science (sp. n.) and report taxonomic changes that affect the species presented in the 2012 catalogue. Nearly three dozen species have been added to the Nearctic Caraboidea (Geadephaga) since Bousquet started the species count at 2678 in 2012.

I (=PWM) look forward to new geographic records (rec. n.) and taxonomic notices obtained from any of the following sources:

(1) A well-documented image on BugGuide or other website that links the imaged specimen to a new record;
(2) A non-imaged specimen that is reliably identified and held in a secure collection with mounted data & determination label. Please submit state/county/province, year, field collector, determiner, repository;
(3) A discovery in the current literature that provides a new record, a new species, or a taxonomic change;
(4) A new record discovered among the photographed specimens ("photo-vouchers") mailed to a carabid specialist for accurate physical determination;
(5) A comment by expert or non-expert that is either posted here or communicated via personal e-mail to PWM.

Please also alert us to additional specimens that would share a regional record already registered here.

Bear in mind that geographic records are based on artificial political areas of widely varying dimensions. Most new records are not surprising as they simply extend the known range to a neighboring state (USA) or province (Canada). In the United States, a range extension that newly spans two or more states is quite unexpected. A specimen linked to a wide geographic extension could merely represent one "accidental" individual (possibly among a few others) that occurred one time in a small tenuous population. When the unexpected does occur, we naturally wonder about the possibility of human error ("status in dubio"). Error can creep in two ways. The more common way is that the species identification was incorrect ("nomen falsum"). The second way is that the wrong collection data was inadvertently attached to a correctly identified specimen in which case its suspected status becomes "locus falsus" or "inscriptio falsa". A more specific designation is "Patria falsa" when a different country is suspected. For example, I am aware of several instances in which authority Fritz Hieke had attached label "Patria falsa: Europa" to certain old Amara specimens in the Milwaukee Public Museum that bear doubtful data from North America. Scientific acceptance depends on our confidence in the source of new information, confidence in the people variously linked to it, and of course the reassurance that comes from actually seeing the specimen (even if just a photograph) and reading the associated collection data.

Please preserve the scientific importance of new geographic records by pin-mounting a colored paper label beneath the specimen that clearly specifies the new state or province record and the year of acknowledgement. A concise example using abbreviations is "WI rec. n. 2017". Useful added information might be data like "series of 6", "BugGuide 1070406".

Current freeze on Tiger Beetle taxonomy: Not yet reflected by this registry nor yet acknowledged by are certain taxonomic revisions of Cicindelini that were proposed in the popular field guide A Field Guide to the Tiger Beetles of the United States and Canada, 2nd ed., 2015 by Pearson et al. Revisions therein were credited to comparative DNA analysis by Duran, DP & Gwiazdowski, RA (2015) who apparently reported their results in an unpublished (not peer reviewed) work entitled “Systematic revision of Nearctic Cicindelini (Coleoptera: Carabidae: Cicindelinae): Re-evaluating Rivalier’s taxonomy”. This state of unsupported taxonomy is discussed in a review of the 2nd edition by M. Jackson in The Coleopterists Bulletin 71(3): 530 - 531, 2017.

Abbreviations & symbols: det =determination; ex(x) =example(s)=specimen(s); leg. =field collector; sp. n. =species new; syn. =synonym; @ =reposited at (institution or individual); → =moved to; % =opinion/observation by; pers. com. =personal communication to PWM; v.i. =vide infra=see below; mm/dd/yy =date > 2012. Institutions are identified by familiar codens in (1) pp 7-10. Frequent contributors who do not post photographs: FFP =Foster F. Purrington. More shortcuts for taxonomists-curators are posted here.

How to search for a particular state or province on this page: hit Ctrl + F, enter the standard two capital letters that identifies the region, under Options select "Match case", all instances on this page will be highlighted. Steps may vary with operating system.

Please alert us to new geographic records that are documented in these post-2012 publications:
2nd edition (2015) of "Tiger Beetles of US & Canada" by Pearson et al.
2nd edition (2013) of "Checklist of Beetles of Canada & Alaska" by Bousquet et al.

Below this "post-2012 Caraboid Registry" is a list of candidate records to consider. (2)(3)(4)(5)(6)(7)(8)(9)(10)(11)(12)(13)(14)

Acupalpus pumilus FL[Gadsden Co: 2000 Kovarik det/@FFP] GA[Oglethorpe: 2004 leg/det/@FFP] IN[Porter Co & Tippecanoe Co: det PWM @FMNH] VT(12)
Agonoleptus rotundicollis KY image
Agonoleptus thoracicus ME image
Agonum aeruginosum OK(6)
Agonum albicrus TX [Wood Co: 1 ex @TAMU det V.Belov v.i.]
Agonum consimilie WY[Johnson Co: 2010 leg. Kondratieff det/@FFP]
Agonum cupreum ME(10)
Agonum cyanopis TX[Jeff Davis Co: 3 exx @TAMU det V.Belov v.i.]
Agonum galvestonicum MA image | WI(8)
Agonum gratiosum NV(14)
Agonum mutatum VA[per C. Harden comment below]
Agonum nigriceps NV(14)
Agonum picicornoides CO[Gunnison Co: 1992 leg. Kondratieff det/@FFP]
Agonum placidum NV(14)
Agonum rufipes RI(13)
Agra oblongopunctata removed from TX (replaced by A. rileyi sp. n. & A. wickhami sp. n.) and removed from North America [Coleopt. Bull. 71(4): 639-651, 2017]
Agra rileyi sp. n. TX [Coleopt. Bull. 71(4): 639-651, 2017]
Agra wickhami sp. n. TX [Coleopt. Bull. 71(4): 639-651, 2017]
Amara aenea KY image
Amara aeneopolita WI image
Amara anthobia ME(10)
Amara angustata OK(6)
Amara apricaria NM[San Juan Co: 1 ♀ det PWM (stylus rules out related New Mexican A. lindrothi Hieke) leg./@J.Marche]
Amara carinata NV(14)
Amara discors AZ [Mohave Co: Pipe Springs NM, 2012 leg. M. Camacho, 1 ȸ ♂ on loan fr NAUF, det PWM]
Amara idahoana NV(14)
Amara ovata NH/VT(12)
Amara sinuosa NV(14)
Amblygnathus mexcinanus OK image
Amblygnathus subtinctus OK image
Amerinus linearis AL[Monroe Co: 12.VI.2016 leg/det/@FFP] OK(6)
Anchonoderus horni Ciski (syn. A. apicalis Horn) new to USA AZ image
Anillinus acutipennis sp. n. TX(7)
Anillinus alleni sp. n. AR[Coleopt. Bull. 71:289-297, 2017]
Anillinus cavicola sp. n. AL(11)
Anillinus comalensis sp. n. TX(7)
Anillinus davidsoni sp. n. AL(11)
Anillinus forthoodensis sp. n. TX(7)
Anillinus hirsutus sp. n. AL(11)
Anillinus inexpectatus sp. n. TN[Annals of Carnegie Museum 82:225-229, 2014]
Anillinus kingi sp. n. AL(11)
Anillinus tombarri sp. n. AL(11)
Anillinus wisemanensis sp. n. TX (7)
Anisodactylus dulcicollis DC(9) MD[Banisteria 44:23-25, 2014] PA image
Anisodactylus haplomus MD[The Maryland Entomologist 7:35-47, 2017]
Anisodactylus harrisii OK(6)
Anisodactylus laetus ME(10)
Anisodactylus melanopus KY image
Apenes lucidula lucidula WI(8) image
Apristus latens NY[Richmond Co.(Mount Loretto Unique Area): 20.VIII.2016 leg. H.Zirlin, det/@PWM]
Apristus pugetanus NV(14)
Apristus subsulcatus OK(6)
Asaphidion curtum curtum PA image
Axinopalpus biplagiatus NV(14)
Axinopalpus fusciceps NV(14)
Badister flavipes flavipes OK(6) VA(2)
Badister grandiceps VA(2)
Badister maculatus MD image
Badister neopulchellus MD image
Badister obtusus NH(12)
Bembidion acutifrons NV(14)
Bembidion anguliferum NV(14)
Bembidion approximatum NV(14)
Bembidion cheyennense UT["Utah 1874", det PWM @UW-IRC]["UT" %D.Maddison pers. com. 02/14/13]
Bembidion diligens WA(5)
Bembidion egens NE[Scotts Bluff Co: 2010 leg. FFP @J.Pretorius det D. Maddison]
Bembidion ephippigerum NV(14)
Bembidion festivum hilare CA[Lassen Co: 2009 leg/@FFP det D.Maddison]
Bembidion fuchsii NV(14)
Bembidion graphicum NE[Morrill Co: 2011 leg/det/@FFP] NV(14) SD[Perkins Co: 2000 leg. J.Maxwell det/@FFP]
Bembidion grapii RI(13)
Bembidion horni CO[Mesa Co: 1988 leg/@FFP det DR Maddison] NV(14)
Bembidion impotens RI image | image | image
Bembidion incrematum NV(14)
Bembidion intermedium ME(10) NV(14) RI(13)
Bembidion lacunarium AZ[Cochise Co: 20.V.2016 leg/det/@FFP]
Bembidion laxatum NV(14)
Bembidion mckinleyi new to USA UT [Sevier Co. leg/@FFP 2000 det DR Maddison "new ssp."]
Bembidion mundum ID[Boise Co & Lewis Co: 2009 leg/@FFP det D.Maddison]
Bembidion mutatum NV(14)
Bembidion nubiculosum NV(14)
Bembidion o. obscurellum NE[Scotts Bluff Co: 2010 leg. FFP @J.Pretorius det FFP]
Bembidion obscuripenne NV(14)
Bembidion obtusangulum NV(14)
Bembidion patruele NM[De Baca Co: 31.V.2016 leg/det/@FFP]
Bembidion praticola VT(12)
Bembidion quadrulum NV(14)
Bembidion recticolle TX[Jeff Davis Co: 2014 leg/det/@FFP]
Bembidion rusticum WV[Maddison, 2008, Annals of Carnegie Museum 77: 148]
Bembidion sejunctum sejunctum ME(10)
Bembidion striola NV(14)
Bembidion umbratum NV(14) TX[Jeff Davis Co: 2014 leg/det/@FFP]
Bembidion versicolor NV(14)
Bembidion versutum MN[Lyon Co: 2008 leg/det/@FFP]
Bembidion viridicolle MA[Nantucket Co: 1995 leg/@FFP det R.Davidson]
Blemus discus IA image
Brachinus americanus RI(13) VA[per C. Harden comment below]
Brachinus cordicollis OK(6)
Brachinus elongatulus NV(14)
Brachinus kansanus TX[Montague Co: 2003 leg/det/@FFP]
Brachinus phaeocerus NV(14) image
Brachinus quadripennis NV(14)
Brachinus tenuicollis RI(13)
Brachinus vulcanoides ME(10)
Bradycellus californicus AZ[Greenlee Co: 28.V.2016 leg/det/@FFP] WY[Crook Co: 13.VII.2016 leg/det/@FFP]
Bradycellus congener NE[Scotts Bluff Co: 2010 leg/det FFP @J.Pretorius]
Bradycellus harpalinus NY[Richmond Co.(Mount Loretto Unique Area): 23.IX.2016 leg. H.Zirlin, det/@PWM] = New to eastern North America
Bradycellus insulsus MA image | image
Bradycellus lecontei IA[Polk C: 2011 leg. D.Veal det/@FFP]
Bradycellus nigriceps RI(13)
Bradycellus nigrinus RI(13)
Bradycellus nitidus NV(14)
Bradycellus rivalis NV(14)
Calathus ingratus RI(13)
Calleida platynoides CO image
Calleida punctata verified TX ["leg. F.Rauterberg before Oct 1897", det G.Noonan & PWM @MPM] RI(13)
Calleida viridipennis NC several records imaged
Calosoma obsoletum WA(5)
Calosoma prominens NV(14)
Calosoma scrutator UT image | image
Calybe laetulaEga laetula LeConte [ZooKeys 430:1-108, 2014] also see
Calybe salleiEga sallei Chevrolat [ZooKeys 430:1-108, 2014] also see
Carabus auratus auratus NB[Coleopt. Bull. 69:264-266, 2015]
Carabus granulatus ME image SK[Coleopt. Bull. 68:700-702, 2014]
Chlaenius erythropus MD image MD[Banisteria 38:71-84, 2011]
Chlaenius obsoletus NV(14)
Chlaenius pennsylvanicus OK(6)
Chlaenius pennsylvanicus blanditus NV(14)
Chlaenius purpureus OK[Comanche Co: 4.VI.2016 leg/det/@FFP]
Cicindela formosa generosa AL[Cincidela 45:1, 2013]
Cicindela longilabris NU[Akimiski Island: Cicindela 2016 48:69-76]
Cicindela pulchra pulchra UT[Cincidela 45:9, 2013]
Cicindela tranquebarica NU[Akimiski Island: Cicindela 2016 48:69-76]
Cicindela trifasciata ascendens IN image
Cicindelidia subgenusgenus : affects 44 taxa listed in Bousquet (2012); see examples here.
Cicindelidia melissa sp. n. AZ [ZooKeys 464:35-47, 2014] | image
Cicindelidia ocellata rectilatera AR[Cicindela 44:49, 2012] KS[image] MS[Cincidela 45:1, 2013]
Cicindelidia sedecimpunctata sedecimpunctata CO [Cincindela 47:71, 2015]
Clinidium apertum allegheniense should be removed from VA(2) and added to WV based on comments by C. Harden below.
Clivina acuducta RI image
Clivina impressefrons CA[Madera Co: 1 ex det V.Belov @TAMU]
Clivina pallida KY image OK(6)
Clivina planicollis AZ[Santa Cruz Co: 1 ex @TAMU det V.Belov v.i.]
Clivina punctulata NV(14)
Clivina rufa MA image WI image
Colliuris caymanensis TX image
Colliuris ludoviciana DE image | image
Coptodera aerata WI(8)
Cyclotrachelus convivus GA[Greene Co: 2013 leg/det/@FFP] VA[per C. Harden comment below]
Cyclotrachelus nonnitens OK(6)
Cyclotrachelus sodalis colossus OK(6)
Cymindis abbreviata → syn. of C. p. platicollis (3)
Cymindis ampliata → syn. of C. p. platicollis (3)
Cymindis arizonensis NV(14)
Cymindis atripennisCymindis platicollis atripennis stat. n. (3)
Cymindis blanda → syn. of C. p. punctigera (3)
Cymindis complanata MS image
Cymindis elegans VA image
Cymindis neglecta RI image
Cymindis obscura → syn. of C. p. platicollis (3)
Cymindis platicollisCymindis p. platicollis stat. n. ND(3) NM(3)
Cymindis punctigeraCymindis p. punctigera subsp. n. (3)
Cymindis rufostigma sp. n. FL (3) image
Cymindis subcarinata → syn. of C. p. punctigera (3)
Cymindis unicolor RI(13)
Diplocheila impressicollis ME(10)
Diplous aterrimus NV(14)
Diplous filicornis NV(14)
Discoderus amoenus TX[Brewster Co: det/@PWM]
Dromius piceus OK(6)
Discoderus aequalis CO[Prowers Co: leg. H.Zirlin, det/@PWM]
Dyschirius erythrocerus VA(2)
Dyschirius sellatus MI[Schoolcraft Co: det PWM @FMNH]
Dyschirius truncatus OK(6)
Elaphropus anceps DC(9)
Elaphropus anthrax NV(14)
Elaphropuis capax TX[Brazoz Co(2017)/Sabine Co(1975): @TAMU det V.Belov v.i.]
Elaphropus conjugens NV(14)
Elaphropus dolosus ME(10) NV(14) VA image
Elaphropus ferrugineus NH(12) WI(8)
Elaphropus granarius TX[Brazos Co/Burleson Co/San Augustine Co: several exx @TAMU det V.Belov v.i.]
Elaphropus conjugens NM image
Elaphropus obesulus TX: det in question image
Elaphropus parvulus MA [new to eastern North America] image
Elaphropus (Tachyura) quadrisignatus (Duftschmid) new to Western Hemisphere NJ[image] VA(9)
Elaphropus vivax RI(13)
Elaphrus californicus OK(6)
Ellipsoptera puritana removed VA per (2)
Galerita lecontei lecontei NV(14) OK(6) UT [Coleopt. Bull. 67(1):56-57, 2013]
Gastrellarius blanchardi TN image
Habroscelimorpha pamphila AL[Cincidela 45:13, 2013]
Harpalus affinis CA[Pan-Pacific Entomologist 90:66-71, 2014] DC[record overlooked in (9)] UT image | image
Harpalus compar MD [MD Biodiversity/det C. Harden]
Harpalus ellipsis NV(14)
Harpalus fuscipalpus NE[Scotts Bluff Co: 2010 leg/det FFP @J.Pretorius]
Harpalus katiae MD image
Harpalus opacipennis OK(6)
Harpalus rubripes VA(9) WI(8)
Helluomorphoides praeustus praeustus RI(13)
Lachnocrepis parallela TX image
Laemostenus complanatus NV(14)
Lebia abdominalis OK(6)
Lebia arizonica [verified? NV image]
Lebia collaris OK(6)
Lebia fuscata OK (6)
Lebia grandis ME(10)
Lebia lobulata KY image
Lebia perita IA[Dallas Co: 2012 leg. E.Freese det/@FFP] NV(14) TX[Jeff Davis Co: 2014 leg/det/@FFP]
Lebia pleuritica OK(6) See challenge to OK record below by V.Belov 19 Sept 2018
Lebia rugatifrons (Chaudoir) new to USA TX image | image
Lebia variegata new to USA AZ image
Lebia 'UASM Lebia 53' (= atypical color morph of Lebia solea?) LA image
Leptotrachelus dorsalis OK(6) MA image | image
Leptotrachelus pallidulus MS["Hancock Co: det R.Davidson" per D.Hildebrandt pers. com. 02/20/13]
Loricera pilicornis pilicornis NV image(14)
Loxandrus cincinnati OK(6)
Loxandrus nr. circulus VA(9)
Loxandrus duryi OK(6)
Loxandrus erraticus OK(6) VA[Banisteria 45:61-62, 2015]
Loxandrus nitidulus OK(6)
Loxandrus parallelus OK(6)
Loxandrus rectus OK(6)
Loxandrus robustus OK(6)
Loxandrus saphyrinus OK(6)
Loxandrus velocipes OK(6)
Microlestes nigrinus verified NM [Santa Fe Co: 2007 leg. R. Delph, det/@ PWM]
Microlestes pusio VA(9)
Mioptachys flavicauda DE image
Myas coracinus ME(10)
Nebria baumanni Kavanaugh sp. n. NV[Proc. Calif. Acad. Sci. 62:441-450 2015]
Nebria brevicollis TX[Smith Co/2015, Brazos Co/2015 & 2017, all @ TAMU, det V.Belov/D.Kavanaugh]
Nebria nivalis gaspesiana NH(12)
Nomius pygmaeus MO[Carter Co, Fremont, 1 ex @TAMU det V.Belov v.i.]
Notiophilus aeneus AR image
Notiobia cyanippa new to USA AZ image | image | image
Notiobia nitidipennis KY image
Notiobia terminata NV(14)
Notiophilus aquaticus NJ image
Notiophilus biguttatus MA images
Notiophilus nitens NV(14)
Oodes americanus DC(9) OK(6)
Oodes brevis WI(8)
Olisthopus brevicornis IN[Porter Co. det/@PWM]
Olisthopus iterans AR[Newton Co. det/@PWM]
Olisthopus micans ME(10)
Omophron labiatum RI image | image
Paraclivina ferrea NE[Scotts Bluff Co: 2010 leg. FFP @J.Pretorius det FFP] VT(12)
Paraclivina striatopunctata MD[Banisteria 38:71-84, 2011]
Paraclivina sulcipennis GA[Wheerler Co: 1♀ 2015 leg/det/@FFP]
Panagaeus fasciatus ME(10)
Pasimachus sublaevis DE image
Patrobus fossifrons NV(14)
Pentagonica bicolor AL image GA[Wheerler Co: 1♀ 2015 leg/det/@FFP]
Pentagonica picticornis ME(10)
Philodes rectangulus NH(12)
Philophuga viridis amoena SD image
Platynus acuminatus new to USA AZ[image Cochise Co: 4 exx leg. C.Melton, det/@PWM] | more images
Platynus opaculus VA(9)
Platynus daviesi NH/VT(12)
Plochionus timidus OK(6) KY image
Polyderis rufotestacea NV(14)
Pseudamara arenaria → tribe Zabrini by Gomez et al. 2016.
Pseudaptinus (Thalpius) dorsalis MO image NC image
Pseudaptinus (Thalpius) rufulus NV(14) WA(5)
Pseudomorpha huachinera sp. n. AZ(4)
Pseudomorpha patagonia sp. n. AZ(4)
Pseudomorpha penablanca sp. n. AZ(4)
Pseudomorpha pima sp. n. AZ(4)
Pseudomorpha santacruz sp. n. AZ(4)
Pseudomorpha santarita sp. n. AZ(4) NM(4)
Pterostichus corrusculus VT(12)
Pterostichus escarinatus CO[Clear Crk. Co: 2014 leg/det/@FFP]
Pterostichus (Anilloferonia) lanei (Hatch, 1935) → replacement Pterostichus (Anilloferonia) diana LaBonte, 2013 [Zootaxa 3682 (4):563-571]
Pterostichus luctuosus NV(14)
Pterostichus permundus DC(9) MD image | The Maryland Entomologist 7:35-47, 2017 VA(9)
Pterostichus punctatissimus WI(8)
Pterostichus sculptus VA(9)
Pterostichus subacutus AR[Coleopt. Bull. 70(2):309-313, 2016]
Pterostichus tenuis VA image
Pterostichus vernalis VT(12)
Rhadine caudata NC[first documented state record claimed by Riley & Browne in Coleopt. Bull. 68(2): 239-240, 2014. NC status given earlier by Bousquet (2012)]
Rhadine nivalis AZ image | image | image
Scaphinotus fissicollis WI image
Scaphinotus hoodooensis sp. n. MT[Proceed. Calif. Acad. Sci. 62:381-396, 2015]
Scarites subterraneus NV(14)
Scarites vicinus DC(9) GA[Cherokee Co: det/@PWM] MD[Banisteria 44:23-25, 2014] VA(9)
Schizogenius depressus NV(14)
Schizogenius lineolatus GA[Greene Co: 2013 leg/det/@FFP]
Selenophorus fabricii sp. n. FL [Zookeys 690: 1-195, 2017]
Selenophorus famulus NV(14)
Selenophorus fatuus KY image
Selenophorus integer removed from FL (replaced in FL by S. fabricii sp. n.)
Selenophorus palliatus verified VA[Sussex Co: 1♂ det PWM leg. R. Whalen @Virginia Polytech Institute]
Selenophorus pedicularius KY image
Sericoda obsoleta OK(6)
Somotrichus unifasciatus verified TX image; VA[Powhatan Co: leg/det/@AV.Evans per C.Harden]
Stenocrepis duodecimstriata RI image | image | image | image
Stenocrepis mexicana MD[The Maryland Entomologist 7:35-47, 2017]
Stenolophus comma NV(14)
Stenolophus fuliginosus NV(14)
Stenolophus lineola WA(5)
Stenolophus maculatus NM[Eddy Co: 2♀ 2014 leg/det/@FFP]
Stenolophus ochropezus NV(14)
Stomis termitiformis CA image
Synuchus impunctatus verified−CO[Animas Co: ♂♀ 2014 leg/det/@FFP] AR image
Tachys (Paratachys) oblitus WI(8)
Tachys (Paratachys) potomaca DC(9)
Tachys (Paratachys) scitulus RI(13)
Tachys (Tachys) mordax TX image
Tachys (Tachys) vittiger TX(v.i.) image
Tachyta angulata OK(6)
Tachyta parvicornis ME(10) OK[Latimer Co: 6 exx 1993-97 @TAMU det V.Belov v.i.]
Tetragonoderus intersectus OK image
Tetraleucus picticornis OK image
Thalpius: see Pseudaptinus
Trachypachus inermis NM image, SD[as syn. T. holmbergi in Coleopt. Bull. 66:357, 2012]
Trechus quadristriatus VA[per C. Harden comment below]
Trechus tenuiscapus NV(14)
Trichotichnus fulgens RI(13) image | image | image
Zuphium punctipenne Bates new to USA TX[Hidalgo Co: det RL Davidson pers. com. 2012 @?TAMU]

These claims for new geographic records require species confirmation in the opinion of PWM.
"Poecilus cursitor" WA image
"Stolonis intercepta" OK(6) and several other species listed 4/5/2014 in bold print are questionably assigned to "OK"(6).

Items removed from the registry.
Bembidion cf. or nr. fortestriatum AZ [UW-IRC: Graham Co, det PWM; Tree of Life website: Greenlee Co, det Maddison 2005] discussed below.

A cited work that affects more than three species is assigned a number as follows.
1. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. Bousquet, Y. 2012. ZooKeys 245:1–1722.

2. New and Additional Records of Ground Beetles and Wrinkled Bark Beetles for Virginia (Coleoptera: Carabidae, Rhysodidae). Roble, S.M. & R.L. Hoffman. 2012. Banisteria No 40:42-48.

3. A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U.S.A. Hunting, W.M. 2013. ZooKeys 259: 1–73.

4. Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group. Erwin, T.L. & L.M. Amundson. 2013. ZooKeys 362:29-54.

5. Ground beetles (Coleoptera, Carabidae) of the Hanford Nuclear Site in south-central Washington State. Looney, C., R.S. Zack, & J.R. LaBonte. 2014. Zookeys 396: 13-42.

6. List of Coleoptera Collected in Latimer County, Oklahoma by Karl Stephan (2002). [Therein the uncertain species cited 4/5/2014 are marked in bold print]

7. Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution. Sokolov, I.M., J.R. Reddell, & D.H. Kavanaugh D.H. 2014. ZooKeys 417:71-101.

8. Additions to the Checklist of Wisconsin Ground Beetles (Coleoptera: Carabidae) including Adventive Harpalus rubripes (Duftschmid) among Seven New State Records. Messer, P.W. 2014. The Great Lakes Entomologist 47: 66-72.

9. Twelve Ground Beetles New to Virginia or the District of Columbia and an Annotated Checklist of the Geadephaga (Coleoptera, Adephaga) from the George Washington Memorial Parkway. Steury, B., & P.W. Messer. 2014. Banisteria 43:40-55.

10. The Ground Beetle (Coleoptera: Carabidae) Fauna of Maine, USA. Dearborn, R.G., R.E. Nelson, C. Donahue, R.T. Bell, & R.P. Webster. 2014. The Coleopterists Bulletin 68: 441 - 599.

11. Five new species of Anillinus Casey from Alabama with a key to the Alabama species (Carabidae: Trechinae: Bembidiini). Sokolov, I.M. 2012. Annals of Carnegie Museum 81:61-71.

12. Carabidae of Vermont and New Hampshire. Bell, R.T. 2015. Shire Press.

13. The Beetle Fauna of Rhode Island. Sikes, D.S. 2004. [NB: Some of the RI records in Sikes' well-documented 2004 list were likely omitted by Bousquet (2012) because they were based on very old specimens identified before modern species keys became available.]

14. Additions to the knowledge of Nevada carabid beetles (Coleoptera: Carabidae) and a preliminary list of carabids from the Great Basin National Park. Will, K., Madan, R., Hsu, H. 2017. Biodiversity Data Journal 5: 1- 23.

Lebia pleuritica OK record challenged
i looked at the specimen (in TAMUIC); the ID is G.E.Ball's, but still the beast has metatarsomere 4 rather modestly emarginate, not bilobed, so it's probably L. tuckeri instead, imho

Tachys mordax in TX

Some new VA and WV records, and a deletion for VA -
In upcoming issue of Banisteria, these are reported (in a paper by myself) as new to VA, (in addition to Cymindis elegans and Somotrichus unifasciatus already reported on this page):
Agonum mutatum
Brachinus americanus
Cyclotrachelus convivus
Elaphropus dolosus
Pterostichus tenuis
Trechus quadristriatus

Photos of these species will be posted soon.

Re-examination of Clinidium apertum allegheniensis in VMNH collection revealed that the specimens had been misidentified, and the species is removed from the VA list in the paper, but is given WV status based on material at CMNH and fresh collections made last year by Bob Davidson, Bob Acciavatti, and myself.

Elaphropus capax in TX
Brazos Co. iii.2017; Sabine Co. vii.1975

Elaphropus granarius in TX confirmed
multiple specimens from the following counties: Angelina, Brazos (incl. the famous Lick Creek Park and my own backyard), Burleson, Fayette, Ft Bend, Milam, Refugio, San Augustine, Walker... à suivre

Tachys vittiger in TX
2 specimens collected in the Brownswille area (Cameron Co.)

hope i'm right about its identity, it's a distinctive tachyine; will ask Terry Erwin about lookalikes

id confirmed by Terry Erwin

Tachyta parvicornis in OK
found a series of six (iii-iv 1993-97) from Latimer Co. in TAMUIC; pretty distinctive beast

Clivina planicollis in AZ, & more TAMUIC stuff
1 ex., Sta Cruz Co.; plus ~40 specimens from several Mexican states (Morelos, NL, Oaxaca, SLP, Tmpls., Veracruz,... country record, i guess); IDs are mine

i also found a specimen from "Madera Co., Calif." that looks exactly like C. impressefrons and keys out snugly to that species, too; the label is reliable, as far as i can tell, and detailed; so after having played with the bug for some time to make sure other options can be ruled out, i decided to call it just that and make it official

Notiophilus aeneus - Arkansas

Agonum cyanopis (Bates): TX
Jeff Davis Co., 3 specimens
again, not surprising

Nomius pygmaeus in MO
found a specimen taken @ uv lights in Fremont Co., MO

not unexpected, of course -- but the bug is too cute and nowhere common

Olisthopus & Agonum albicrus (Dej.) in TX
just found in TAMUIC 3 specimens of Olisthopus (similar to micans) from e. TX (Tyler & Sabine counties) and one Agonum albicrus from Wood Co.

Selenophorus fatuus - Kentucky

Harpalus katiae - Maryland

Clivina pallida Say - KY

Selenophorus pedicularius Dejean - KY

Calleida viridipennis (Say)
Several images in the guide from North Carolina. NC is not recorded in Bousquet 2012.

Pasimachus sublaevis new for Delaware


potential KS record

Trachypachus inermis for NM

Gastrellarius blanchardi, TN


Omophron labiatum, RI
I'm pretty sure this is right, but I'd appreciate confirmation. I found three specimens that I believe to be this species (and I've photographed and collected two of them). Info page says "pretty distinctive" but I'm hardly an expert. I also found what I believe is an O. americanum and very many O. tesselatum (currently the only two species known from RI) along the same stretch of shoreline. All images are at family level.

Somotrichus unifasciatus, Virginia
Based on a specimen in the Art. V. Evans collection (AVEC). Collected from a UV light trap set at Powhatan State park in June 2015. Det. by Art Evans, who also notified Yves Bousquet about the record.

Agonoleptus thoracicus, Maine

(the ventral view looks right for that species, to my eye)

Cymindis elegans, Virginia

I've notified Yves Bousquet about it.

Stomis termitiformis, CA

I can vouch for Calosoma scru
I can vouch for Calosoma scrutator being well established in Washington County, Utah. I have seen dozens of specimens from multiple collectors in two different collections. The most common localities are Lytle Preserve and Beaver Dam Wash. Neither Erwin's or Bousquet's catalogs recognise it from Utah.

Thank you for candidate Calosoma scrutator in UT.
While I personally don't doubt this claim, C. scrutator cannot be posted as a new record until I learn of at least one specific specimen (= voucher) that is mounted with compete data label and a determination label (species name followed by name of the determiner). As the guidelines say above, I need at least county, determiner, and repository. However, it would be best if you or another collector could get a Utahn Calosoma scrutator imaged on BugGuide.

sorry it took so long
I added two records under C. scrutator, i would choose the specimen collected by Baumann and Clark, out of the two submissions, to be the first representative on this list.

No problem. Give me a week or
No problem. Give me a week or so to re-visit the specimens.

Agonoleptus rotundicollis (Haldeman), KY

Just a note, maybe is need an update on above list.
Colliuris tetrastigma on BG page is under Colliuris caymanensis new name.
(Colliuris tetrastigma caymanensis Darlington; elevated to species by Bousquet, 2010)

Colliuris caymanensis Darlington
is indeed the valid name acknowledged on page 1270 of the catalogue by Bousquet (2012). Thanks Iustin for alerting us to the new taxonomic combination. 

new to continent?

Lebia lobulata LeConte, KY

Anisodactylus melanopus, KY

Notiobia nitidipennis
new for KY:

Chlaenius erythropus MD

Amara aenea, KY
Didn't see KY listed in Bousquet, though surrounding states are all represented.

B. fortestriatum is probably not in AZ
Those specimens of "Bembidion fortestriatum" that are mentioned from Greenlee Co. are almost certainly not B. fortestriatum, even though there is a claim in this list that I identified them. There is a species in Greenlee County that is similar to B. fortestriatum, which I typically call "B. cf fortestriatum" or "B. sp. nr. fortestriatum", but it is not B. fortestriatum.

New list of removed species is started
thanks to your clarification David. Sorry in this case for not notating your qualifiers "cf." or "nr." in my original hand-written database. I'll look for those labels again when I revisit collections at UW-Madison-IRC and Milwaukee Public Museum later this year. For comparison I have a good example of B. fortestriatum that I recently captured in WI, Ashland Co. This is just the kind of expert feed back I was hoping for!

The designation "Bembidion fortestriatum" in AZ, Greenlee Co,
presented without the qualifier "cf." or "nr." in the name, was not derived from a curated/labeled specimen as I mistakenly suggested above. I now recall that the source was the information posted on Tree of Life website. It shows beautiful habitus images of the beetle culprit by determiner/photographer David Maddison, 2005. The straight forward name "Bembidion fortestriatum" that I assigned to a single examined specimen from Arizona-Graham County (adjacent to Greenlee Co) had apparently been influenced by my reading the same designation posted on that website. David and I had a good telephone discussion about the matter today. He intends to make the necessary change to the species name on the website soon.

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