Keywords: Caraboidea, Carabidae, carabid, Tiger Beetles, new species, North America, United States, Canada
*Frequent updates without notice*
This website is an interactive clearinghouse where users of BugGuide.net informally provide new information to supplement the geographic distributions, taxonomic classification, and scientific names published in the massive 2012 catalogue of Nearctic Geadephaga by Yves Bousquet
(1) [YB2012].
NB: Not all photographed specimens in this list have had their species identifications corroborated physically by an authority.
The large beetle group designated "Geadephaga", commonly known as the "Ground Beetles", is the superfamily Caraboidea composed of the three families
Trachypachidae,
Rhysodidae, and
Carabidae. The latter family is by far the largest one. Its members, informally named "carabids", account for 99.55 percent of the roughly 2700 known species in the
Nearctic Caraboidea north of Mexico. It is this faunal region that concerns BugGuide and this
post-2012 Caraboidea Registry/Forum. The family Carabidae includes several subfamilies of which Cicindelinae is represented by the popular "Tiger Beetles". See the comments below regarding the state of Tiger Beetle taxonomy.
Newly documented states, provinces, or territories are entered by their standard abbreviations after the alphabetically-listed species names below. In addition, geographic records that confirm the questionable ones enclosed in square brackets by Bousquet, will be annotated here as
verified. Currently 16 species are newly reported to have one of these major range extensions:
new to USA,
new to North America,
new to Western Hemisphere. Enigmatic specimens that might be either undescribed or new to the Nearctic north of Mexico are annotated
unresolved.
In tracking the taxonomic literature, we will include Nearctic species (
sp. n.) and subspecies (
ssp. n.) that are new to science and report taxonomic changes that affect the species presented in the 2012 catalogue. The numerous recent nomenclatural changes that affect the Tiger Beetles in Bousquet (2012) are not yet incorporated at this registry. About five dozen (50 in 2022) new species have been added to the Nearctic Caraboidea (Geadephaga) since Bousquet started the species count at
2678 in 2012. Also affecting the count are several cited species that have been either removed from synonymy or moved to synonymy.
I (=
PWM) look forward to new geographic records (rec. n.) and taxonomic notices obtained from any of the following sources:
(1) A well-documented image on
BugGuide,
iNaturalist, or other website that links the imaged specimen to a new record;
(2) A non-imaged specimen that is reliably identified and held in a secure collection with mounted data & determination label.
Please submit state/county/province, date, determiner, repository;
(3) A discovery in the current literature that provides a new record, a new species, or a taxonomic change;
(4) A new record discovered among the photographed specimens ("photo-vouchers") mailed to a carabid specialist for accurate physical determination;
(5) A comment by expert or non-expert that is either posted here or communicated via personal e-mail to PWM.
Please also alert us to additional specimens that would
share a regional record already registered here. Rather than posting my feedback or questions on this page, I usually contact the submitter via the image source page or via personal e-mail.
Please bear in mind that geographic records are based on artificial political areas of widely varying dimensions. Most new records are not surprising as they simply extend the known range to a neighboring political region. In the United States, for example, a range extension that newly spans two or more states is quite unexpected. A specimen linked to a wide geographic extension could merely represent one "accidental" individual (possibly among a few others) that occurred one time in a small tenuous population. When the unexpected does occur, we naturally wonder about the possibility of human error ("status in dubio"). Error can creep in two ways. The more common way is that the species identification was incorrect ("nomen falsum"). The second way is that the wrong collection data was inadvertently attached to a correctly identified specimen in which case its suspected status becomes "locus falsus" or "inscriptio falsa". A more specific designation is "Patria falsa" when a different country is suspected. For example, I am aware of several instances in which authority Fritz Hieke had attached label "Patria falsa: Europa" to certain old
Amara specimens in the Milwaukee Public Museum that bear doubtful data from North America. Scientific acceptance depends on our confidence in the source of new information, confidence in the people variously linked to it, and of course the reassurance that comes from actually seeing the specimen (even if just a photograph) and reading the associated collection data.
Please preserve the scientific importance of new geographic records by pin-mounting a colored paper label beneath the specimen that clearly specifies the new state or province record and the year of acknowledgement. A concise example using abbreviations is "WI rec. n. 2017". Useful added information might be data like "series of 6", "BugGuide 1070406".
Discussions on the taxonomic state of Nearctic Tiger Beetles.
Pre-2019 statements: Not yet reflected by this registry nor yet acknowledged by BugGuide.net are certain taxonomic revisions of Cicindelini that were proposed in the popular field guide
A Field Guide to the Tiger Beetles of the United States and Canada, 2nd ed., 2015 by Pearson et al. Revisions therein were credited to comparative DNA analysis by Duran, DP & Gwiazdowski, RA (2015) who apparently reported their results in an unpublished (not peer reviewed) work entitled “Systematic revision of Nearctic Cicindelini (Coleoptera: Carabidae: Cicindelinae): Re-evaluating Rivalier’s taxonomy”. This state of unsupported taxonomy is discussed in a review of the 2nd edition by M. Jackson in The Coleopterists Bulletin 71(3): 530 - 531, 2017.
July 15, 2019: See the news below from V. Belov stating that papers for fixing the problematical taxonomy have been submitted.
September, 2019: A paper by Duran & Gough in
Insecta Munda Sept 27, 2019 (0727): 1-12 clears the way for the proposed nomenclatural changes. See the developing thread below.
NB: The new Tiger Beetle names that affect Bousquet (2012) are now successfully incorporated into BugGuide.net, but the changes are not yet listed in this registry. It is unresolved whether BugGuide will follow the family status of tiger beetles as advocated in a paper by Duran & Gough (2020). See ongoing discussions
here.
Abbreviations & symbols: ♂ ♀ sex; det =determination; ex(x) =example(s)=specimen(s); leg. =field collector; sp.(ssp.) n. =species (subspecies) new; syn. =synonym; @ =reposited at (institution or individual); → =moved to sr. synonym or new combination; % =opinion/observation by; pers. com. =personal communication to PWM; tbd =to be determined; v.i. =vide infra=see below (comment); mm/dd/yy =date > 2012. Institutions are identified by familiar acronyms in
(1) pp 7-10.
Frequent contributors: FFP =Foster F. Purrington
†2023; CWH =Curt W. Harden.
How to search for a particular state, province, or territory on this page: hit Ctrl + F; enter the standard two capital letters that identifies the region; under Options select "Match case" and "Match whole word"; all instances on this page will be highlighted.
Please alert us to new geographic records that are documented in these post-2012 publications which I have not yet reviewed:
2nd edition (2015) of "Tiger Beetles of US & Canada" by Pearson et al.
2nd edition (2013) of "Checklist of Beetles of Canada & Alaska" by Bousquet et al.
Below this post-2012 Caraboidea Registry is a list of candidate records to consider. (2)(3)(4)(5)(6)(7)(8)(9)(10)(11)(12)(13)(14)(15)(16)(17)(18)(19)(20)(21)(22)(23)(24)(25)(26)(27)
Acupalpus partiarius ND
image NM [Quay Co, Roosevelt Co, The Coleopterists Bulletin 74(1):21, 2020]
Acupalpus pumilus FL[Gadsden Co: 2000 Kovarik det/@FFP] GA[Oglethorpe: 2004 leg/det/@FFP] IN[Porter Co & Tippecanoe Co: det PWM @FMNH] NB
(24) VT
(12)
Acupalpus tener NV
image
Agonoleptus convexulus new to USA FL
image
Agonoleptus rotundicollis KY
image
Agonoleptus thoracicus ME
image WV
image
Agonum aeruginosum OK
(6)
Agonum albicrus TX [Wood Co: 1 ex @TAMU det V.Belov v.i.]
Agonum consimilie WY[Johnson Co: 2010 leg. Kondratieff det/@FFP]
Agonum cupreum ME
(10)
Agonum cyanopis TX[Jeff Davis Co: 3 exx @TAMU det V.Belov v.i.]
Agonum galvestonicum MA
image | MD
image | WI
(8)
Agonum gratiosum NV
(14)
Agonum harrisii CA
image image image
Agonum mutatum VA
(15)
Agonum nigriceps NV
(14)
Agonum picicornoides CO[Gunnison Co: 1992 leg. Kondratieff det/@FFP]
Agonum placidum NV
(14)
Agonum propinquum UT
image
Agonum rufipes RI
(13)
Agonum tenue CO [1♂ Buena Vista [Chaffee Co.], leg. H.F. Wickham Jul 1-6 1896, det PWM 2022, @PWM or WIRC]
Agra oblongopunctata removed from TX (replaced by
A. rileyi sp. n. &
A. wickhami sp. n.) and
removed from North America [Coleopt. Bull. 71(4): 639-651, 2017]
Agra rileyi sp. n. TX [Coleopt. Bull. 71(4): 639-651, 2017]
image
Agra wickhami sp. n. TX [Coleopt. Bull. 71(4): 639-651, 2017]
Amara aenea KY
image ND
image NM[Bernalillo Co., 1♀ leg. RJ Buss 1992, det/@PWM] TX[Madison Co., 1♂ leg. TR Clifton 2017, @BT Raber, det PWM]
Amara aeneopolita WI
image
Amara anthobia ME
(10)
Amara angustata OK
(6)
Amara apachensis OK [Cimarron Co. in western OK, 1♀ leg. RJ Buss 2009, det/@PWM 2021]
Amara apricaria NM[San Juan Co: 1♀ det PWM (stylus rules out related New Mexican
A. lindrothi Hieke) leg./@J.Marche]
Amara aurata recheck against A. convexa: NM[Otero Co. in southern NM, 1♂ leg. RJ Buss 2006, det/@PWM 2021]
Amara avida DE
image
Amara c. californica KS[Morton Co: 1♂ 2015-X-15, leg. W.Chatfield-Taylor, det/@PWM]
Amara carinata NV
(14)
Amara discors AZ [Mohave Co: Pipe Springs NM, 2012 leg. M. Camacho, 1 ȸ ♂ on loan fr NAUF, det PWM] TX[Jeff Davis Co., 10 exx V-VI 2020, leg.Raber/Heffern, det PWM, @Raber/PWM]
Amara flebilis NC
image
Amara idahoana NV
(14)
Amara latior OK [Cimarron Co., 1♀ RJ Buss 2009, det/@ PWM 2021]
Amara lunicollis ND
image
Amara musculis TX
confirmed [Lamar Co., 1♀ 2003, leg.Godwin, @BTRaber, det PWM]
Amara ovata NH/VT
(12)
Amara sinuosa NV
(14)
Amara torrida SD[Pennington Co., 1♂ 1989-VIII-4, leg. RJ Buss, det/@ PWM 2021]
Amblycheila katzi sp. n. TX
image [reference given under species Info tab]
Amblygnathus balli sp. n. AZ (see
Shpeley & Van Devender, 2021)
Amblygnathus interior new to USA AZ
images
Amblygnathus mexicanus OK
image
Amblygnathus subtinctus OK
image
Amerinus linearis AL[Monroe Co: 12.VI.2016 leg/det/@FFP] OK
(6)
Amphasia interstitialis ND
image
Amphasia sericea ND
image
Anatrichis minuta NJ
image
Anchomenus aeneolus NV
image
Anchonoderus horni Ciski (syn.
A. apicalis Horn)
new to USA: AZ
image |
image
Anillinus acutipennis sp. n.(7) TX
Anillinus albrittonorum sp. n. FL [
Sokolov & Schnepp, 2021]
Anillinus alleni sp. n. AR[Coleopt. Bull. 71:289-297, 2017]
Anillinus arenicollis sp. n.(11) SC
image
Anillinus bexarensis sp. n. TX[Sokolov 2022, DOI: 10.3897/subtbiol.44.91002]
Anillinus castaneus sp. n.(27) SC
Anillinus cavicola sp. n. AL
(11)
Anillinus choestoea sp. n.(27) SC
image
Anillinus cherokee GA
image SC [Sokolov & Carlton 2010: p. 15 - v.i.]
Anillinus clinei sp. n. AL
(19)
Anillinus comalensis sp. n. TX
(7)
Anillinus davidsoni sp. n. AL
(11)
Anillinus dentatus sp. n.(27) SC
image
Anillinus elongatus removed: VA
image
Anillus felicianus sp. n. LA [Sokolov, 2021, Zookeys 1016:63-76]
Anillinus folkertsioides sp. n. AL
(19)
Anillinus forthoodensis sp. n. TX
(7)
Anillinus hildebrandti sp. n. AL
(19)
Anillinus hirsutus sp. n. AL
(11)
Anillinus humicolus sp. n. AL
(19)
Anillinus inexpectatus sp. n. TN[
Annals of Carnegie Museum 82:225-229, 2014]
Anillinus jancae sp. n.(27) SC
image
Anillinus kingi sp. n.(11) NC SC
Anillinus mica sp. n.(27) SC
image
Anillinus micamicus sp. n.(27) SC
image
Anillinus montrex sp. n.(26) NC SC
image
Anillinus murrayae SC
image
Anillinus pecki VA & TN
(26)
Anillinus pittsylvanicus sp. n.(26) VA
image
Anillinus reddelli sp. n. TX[Sokolov 2022, DOI: 10.3897/subtbiol.44.91002]
Anillus relictus sp. n. AL [Sokolov, 2021, Zookeys 1016:63-76]
Anillinus seneca sp. n.(27) SC
image
Anillinus simplex sp. n.(27) SC
Anillinus tombarri sp. n. AL
(11)
Anillinus uwharrie sp. n.(26) NC
image
Anillinus wisemanensis sp. n. TX
(7)
Anisocnemus amblygonus "AZ" likely
Patria falsa image
Anisodactlylus binotatus ID
image UT
image
Anisodactylus dulcicollis DC
(9) MD[
Banisteria 44:23-25, 2014] PA
image
Anisodactylus furvus DC
image
Anisodactylus haplomus MD[
The Maryland Entomologist 7:35-47, 2017]
Anisodactylus harrisii OK
(6)
Anisodactylus laetus ME
(10)
Anisodactylus melanopus KY
image
Anisodactylus merula NB
(24)
Anisodactylus nigrita UT
image WY
image
Anisodactylus ovularis ND
image
Anisodactylus porosus UT
image
Anisodactylus verticalis AL [Monroe Co., Jun 2016, 3 exx @FSCA, det PWM]
Apenes lucidula lucidula WI
(8) image
Apristus latens NY[Richmond Co.(Mount Loretto Unique Area): 20.VIII.2016 leg. H.Zirlin, det/@PWM] MO
image
Apristus pugetanus NV
(14)
Apristus subsulcatus OK
(6)
Ardistomis schaumii MO
image
Asaphidion curtum curtum PA
image
Aspidoglossa subangulata LA [West Feliciana Parish, 13-14 IV 2014, leg. MH Evans, det/@PWM]
Axinopalpus biplagiatus NV
(14)
Axinopalpus illectus OR
image image image
Axinopalpus fusciceps NV
(14)
Badister flavipes flavipes OK
(6) VA
(2)
Badister grandiceps VA
(2)
Badister maculatus MD
image
Badister neopulchellus MD
image
Badister obtusus NH
(12) OR
image
Badister transversus TX[San Patricio Co: 2013, det V.Belov v.i. awaiting image]
Bembidion acutifrons NV
(14)
Bembidion (Neja) ambiguum new to North America CA [
PDF]
Bembidion amnicum LA
image
Bembidion anguliferum NV
(14)
Bembidion approximatum NV
(14)
Bemdidion brownorum sp. n. CA
[ZooKeys 1156: 87-106 (2023)]
Bembidion (Liocosmius) cachagua sp. n. CA
LaBonte & Madison 2023: p433
Bembidion cheyennense UT["Utah 1874", det PWM @UW-IRC]["UT" %D.Maddison pers. com. 02/14/13][
image]
Bembidion concretum ID
image
Bembidion cooperi sp. n. AZ
(21), NM
(21)
Bembidion corgenoma sp. n. BC to several western states [Maddison, 2020, Zookeys 1007: 85-128] CA
image
Bembidion darlingtonielum sp. n. CA
(21)
Bembidion diligens WA
(5)
Bembidion egens NE[Scotts Bluff Co: 2010 leg. FFP @J.Pretorius det D. Maddison]
Bembidion ephippigerum NV
(14)
Bembidion festivum hilare CA[Lassen Co: 2009 leg/@FFP det D.Maddison]
(21)
Bembidion fuchsii NV
(14)
Bembidion graphicum NE[Morrill Co: 2011 leg/det/@FFP] NV
(14) SD[Perkins Co: 2000 leg. J.Maxwell det/@FFP]
Bembidion grapii RI
(13)
Bembidion horni CO[Mesa Co: 1988 leg/@FFP det DR Maddison]
(21) NV
(14)
Bembidion impotens OR? [v.i. CEJG comment 19 Dec 2023] RI
image |
image |
image
Bembidion incrematum NV
(14) UT
image
Bembidion intermedium ME
(10) NV
(14) RI
(13)
Bembidion lacunarium AZ[Cochise Co: 20.V.2016 leg/det/@FFP]
Bembidion laxatum NV
(14)
Bembidion mckinleyi new to USA UT [Sevier Co. leg/@FFP 2000 det DR Maddison "new ssp."]
Bembidion mexicanum UT
image
Bembidion mimbres sp. n. AZ, NM [Maddison, 2020, Zookeys 1007: 85-128]
Bembidion mundum ID[Boise Co & Lewis Co: 2009 leg/@FFP det D.Maddison]
(21)
Bembidion mutatum NV
(14)
Bembidion nubiculosum NV
(14)
Bembidion o. obscurellum NE[Scotts Bluff Co: 2010 leg. FFP @J.Pretorius det FFP]
Bembidion obscuripenne NV
(14)
Bembidion obtusangulum NV
(14)
Bembidion orion sp. n. CA
(21)
Bembidion patruele NM[De Baca Co: 31.V.2016 leg/det/@FFP] UT
image
Bembidion pedicellatum SD
image
Bembidion plagiatum SC
image
Bembidion poculare TX[El Paso Co., leg. N.Brown 2005, @TAMU, det V.Belov (v.i)]
Bembidion praticola VT
(12)
Bembidion quadrulum NV
(14)
Bembidion rapidum OR
image image image
Bembidion recticolle TX[Jeff Davis Co: 2014 leg/det/@FFP]
Bembidion rusticum WV[Maddison, 2008, Annals of Carnegie Museum 77: 148]
Bembidion sejunctum sejunctum ME
(10)
Bembidion striola NV
(14)
Bembidion texanum ON
image
Bembidion umbratum NV
(14) TX[Jeff Davis Co: 2014 leg/det/@FFP]
Bembidion versicolor NV
(14)
Bembidion versutum MN[Lyon Co: 2008 leg/det/@FFP]
Bembidion viridicolle MA[Nantucket Co: 1995 leg/@FFP det R.Davidson] MI[Midland Co, 1944, 2 exx; Montcalm Co, 1942, 1 ex; all @MSU/det PWM]
Blemus discus IA
image
Blethisa julii NU
(16)
Blethisa quadricollis NU
(16)
Brachinus americanus RI
(13) VA
(15)
Brachinus cordicollis OK
(6)
Brachinus elongatulus NV
(14)
Brachinus kansanus TX[Montague Co: 2003 leg/det/@FFP]
Brachinus phaeocerus NV
(14) another NV record removed
Brachinus quadripennis NV
(14)
Brachinus tenuicollis RI
(13)
Brachinus vulcanoides ME
(10)
Bradycellus atrimedeus ND
image
Bradycellus californicus AZ[Greenlee Co: 28.V.2016 leg/det/@FFP] UT[Utah Co: @ TAMU/det V.Belov v.i.
image; more exx det/@ J. Quist v.i.] WY[Crook Co: 13.VII.2016 leg/det/@FFP]
Bradycellus congener NE[Scotts Bluff Co: 2010 leg/det FFP @J.Pretorius]
Bradycellus festinans LA[Rapides Parish: Alexandria, 2.VIII.1970, 1♂ det V.Belov/PWM, @ TAMU] LA[St. Gabriel, iNaturalist 156331628]
Bradycellus harpalinus NY[Richmond Co.(Mount Loretto Unique Area): 23.IX.2016 leg. H.Zirlin, det/@PWM] =
New to eastern North America
Bradycellus insulsus MA
image |
image
Bradycellus lecontei IA[Polk C: 2011 leg. D.Veal det/@FFP]
Bradycellus nigriceps RI
(13)
Bradycellus nigrinus RI
(13)
Bradycellus nitidus NV
(14)
Bradycellus rivalis NV
(14)
Brasiella viridisticta arizonensis NM [
CICINDELA 55(3):77-81, 2023]
Calathus ingratus RI
(13)
Calleida decora MO
image
Calleida platynoides CO
image
Calleida punctata verified TX ["leg. F.Rauterberg before Oct 1897", det G.Noonan & PWM @MPM] RI
(13)
Calleida punctulata AZ
tentative: image image image
Calleida rubricollis new to USA from West Indies: FL
image
Calleida viridipennis AZ
confirmed image NC
several records imaged
Calosoma (Chrysostigma) ampliator new to USA: AZ(Pima Co.)
Calosoma frigidum NT
image
Calosoma obsoletum WA
(5)
Calosoma prominens NV
(14)
Calosoma scrutator UT
image |
image
Calybe laetula →
Ega laetula LeConte [ZooKeys 430:1-108, 2014]
also see
Calybe sallei →
Ega sallei Chevrolat [ZooKeys 430:1-108, 2014]
also see
Carabus auratus auratus NB[
Coleopt. Bull. 69:264-266, 2015]
Carabus granulatus ME
image ND
image SK[
Coleopt. Bull. 68:700-702, 2014]
Carabus nemoralis CO
records
Chlaenius brevilabris QC
image
Chlaenius erythropus MD
image MD[
Banisteria 38:71-84, 2011]
Chlaenius obsoletus NV
(14)
Chlaenius pennsylvanicus OK
(6)
Chlaenius pennsylvanicus blanditus NV
(14)
Chlaenius purpureus OK[Comanche Co: 4.VI.2016 leg/det/@FFP]
Chlaenius tomentosus NB
(24)
Cicindela formosa gaumeri ssp. n. northwestern CO
French, Bell & Acorn 2020
Cicindela formosa generosa AL[
CICINDELA 45:1, 2013]
Cicindela limbata nympha WY[Carbon Co: several, 2019, leg/det/@ML Brust & @PWMC]
Cicindela longilabris NU[Akimiski Island:
CICINDELA 48:69-76, 2016]
Cicindela pulchra pulchra UT[
CICINDELA 45:9, 2013]
Cicindela scutellaris NB
(24)
Cicindela sexguttata WY [
CICINDELA 52(4): 86, 2020]
Cicindela tranquebarica NU[Akimiski Island:
CICINDELA 48:69-76, 2016] YT
(24)
Cicindela tranquebarica admiscens Casey, 1913
new status NM, AZ, UT [CICINDELA 55(2):25, 2023]
Cicindela tranquebarica diffracta Casey, 1909
new synonymy with
C. t. kirbyi CO, NM [CICINDELA 55(2):25, 2023]
Cicindela tranquebarica kirbyi WY [
CICINDELA 52(4): 86, 2020]
Cicindela trifasciata ascendens IN
image
Cicindelidia subgenus →
genus : affects 44 taxa listed in Bousquet (2012); see examples here.
Cicindelidia melissa sp. n. AZ [
ZooKeys 464:35-47, 2014] |
image
Cicindelidia ocellata ocellata verified TX [Bailey Co, The Coleopterists Bulletin 74(1):19, 2020]
Cicindelidia ocellata rectilatera AR[
CICINDELA 44:49, 2012] KS[
image] MS[
CICINDELA 45:1, 2013]
Cicindelidia punctulata AZ[Yuma Co.,
CICINDELA 2019 51:23] CA[Inyo Co.,
CICINDELA 2019 51:23] NV[Nye Co., Elko Co.,
CICINDELA 2019 51:23] OR[Ontario,
CICINDELA 2024 56(1):13]
Cicindelidia sedecimpunctata sedecimpunctata CO [
CICINDELA 47:71, 2015]
Clinidium apertum allegheniense should be removed from VA
(2) and added to WV
(15):
image |
image
Clivina acuducta RI
image
Clivina collaris OR
image
Clivina fossor ND
image
Clivina impressefrons CA[Madera Co: 1 ex det V.Belov @TAMU]
Clivina pallida KY
image OK
(6)
Clivina planicollis AZ[Santa Cruz Co: 1 ex @TAMU det V.Belov v.i.]
Clivina punctulata NV
(14)
Clivina rufa MA
image WI
image
Colliuris caymanensis TX
image |
image
Colliuris ludoviciana DE
image |
image TX
image
Colliuris pensylvanica NB
(24)
Coptodera aerata WI
(8)
Cyclotrachelus convivus GA[Greene Co: 2013 leg/det/@FFP] VA
(15)
Cyclotrachelus fuscatus IN
image VA
image
Cyclotrachelus nonnitens OK
(6)
Cyclotrachelus sodalis colossus OK
(6)
Cyclotrachelus sodalis sodalis MA
image
Cymindis abbreviata → syn. of
C. p. platicollis (3)
Cymindis ampliata → syn. of
C. p. platicollis (3)
Cymindis apache sp. n. AZ [larger specimens labeled "
C. punctifera" are likely
C. apache; see
Dugesiana 31(1): 19-74;
image]
Cymindis arizonensis NV
(14)
Cymindis atripennis →
Cymindis platicollis atripennis stat. n.
(3)
Cymindis blanda → syn. of
C. p. punctigera (3)
Cymindis complanata MS
image
Cymindis elegans VA
image(15)
Cymindis neglecta RI
image
Cymindis obscura → syn. of
C. p. platicollis (3)
Cymindis platicollis →
Cymindis p. platicollis stat. n. NB
(24) ND
(3) NM
(3)
Cymindis p. platicollis CO [new senior synonym
(3) of Coloradan
C. abbreviata &
C. ampliata]
Cymindis punctigera →
Cymindis p. punctigera subsp. n.
(3)
Cymindis rufostigma sp. n. FL
(3) image
Cymindis subcarinata → syn. of
C. p. punctigera (3)
Cymindis unicolor RI
(13)
Diacheila arctica NU
(16) SK
image
Diplocheila impressicollis ME
(10)
Diplocheila obtusa UT
image
Diplous aterrimus NV
(14)
Diplous filicornis NV
(14)
Discoderus aequalis CO[Prowers Co: leg. H.Zirlin, det/@PWM] KS
image |
image
Discoderus amoenus TX[Brewster Co: det/@PWM]
Dromius piceus OK
(6)
Dyschirius abbreviatus SC
image
Dyschirius affinis MI[Macomb Co, 1963, 1 ex @MSU, det G.Parsons & PWM]
Dyschirius erythrocerus VA
(2)
Dyschirius larochellei ON [northern Ontario, 2021, https://doi.org/10.1080/11956860.2021.1872265]
Dyschirius sellatus MI[Schoolcraft Co: det PWM @FMNH]
Dyschirius truncatus OK
(6)
Ega sallei NC[Carteret Co., Coleopterists Bull. 76(3):379] TN
image TN [Polk Co., 22-V-2021, det/@C.Harden v.i.]
Elaphropus anceps DC
(9)
Elaphropus anthrax NV
(14)
Elaphropus capax MI[Midland Co, 1937, 1 ex @MSU, det W.Ruesink & PWM] TX[Brazoz Co(2017)/Sabine Co(1975): @TAMU det V.Belov v.i.]
Elaphropus conjugens NV
(14)
Elaphropus dolosus ME
(10) NV
(14) VA
image(15)
Elaphropus ferrugineus NH
(12) WI
(8)
Elaphropus granarius TX[Brazos Co/Burleson Co/San Augustine Co: several exx @TAMU det V.Belov v.i.]
Elaphropus conjugens NM
image
Elaphropus incurvus VA:
unconfirmed as ex lost image
Elaphropus obesulus TX:
det in question image
Elaphropus (Tachyura) lutea new to North America TN & VA [Coleop. Bull. 78(3): 408-409, 2024] MO[Columbia, 8/2024, leg/det/@CWH, v.i.]
Elaphropus (Tachyura) parvulus new to eastern North America MA
image
Elaphropus (Tachyura) quadrisignatus (Duftschmid)
new to Western Hemisphere NJ[
image] VA
(9)
Elaphropus vivax RI
(13)
Elaphrus americanus NU
(16)
Elaphrus californicus OK
(6)
Elaphrus clairvillei NU
(16)
Elaphrus fuliginosus NU
(16)
Elaphrus lecontei NU
(16)
Ellipsoptera nevadica olmosa removed from AZ, NM[
CICINDELA 54:21, 2022]
Ellipsoptera nevadica playa ssp. n. NM [
CICINDELA 54:21, 2022]
Ellipsoptera puritana removed from VA
(2)
Ellipsopetra rubicunda (Harris, 1911) →
E. marutha Dow, 1911 according to Pearson, Duran, et al.
Eucaerus varicornis NJ
image
Galerita lecontei lecontei NV
(14) OK
(6) UT [Coleopt. Bull. 67(1):56-57, 2013]
Gastrellarius blanchardi TN
image
Habroscelimorpha pamphila AL[
CICINDELA45:13, 2013]
Harpalus:
revised classification of subgenera by Kataev 2023
(25)
Harpalus affinis CA[Pan-Pacific Entomologist 90:66-71, 2014] CO
? image DC[record overlooked in
(9)] UT
image |
image
Harpalus compar MD [
MD Biodiversity/det C. Harden] |
image
Harpalus ellipsis NV
(14)
Harpalus faunus NM [Bernalillo Co., 1986, 1♀ leg./@ RJ Buss, det PWM]
Harpalus fraternus ND
image
Harpalus fulvilabris UT
image WY [Park Co., 1♂ leg. RJ Buss 1995, det/@PWM]
Harpalus fuscipalpus NE[Scotts Bluff Co: 2010 leg/det FFP @J.Pretorius]
Harpalus gravis KS
image
Harpalus katiae MD
image(15)
Harpalus laticeps UT
image
Harpalus opacipennis OK
(6)
Harpalus rubripes IL [Chicago, The Coleop. Bull. 74: 894, 2020] IN
image VA
(9) WA
image WI
(8)
Helluomorphoides praeustus praeustus RI
(13)
Horologion hubbardi sp. n. VA [
image, Subterranean Biology 48: 1–49.
https://doi.org/10.3897/subtbiol.48.114404]
Hybothecus flohri TX [Maverick Co., 18 VIII 2016, 1♀ leg. K.Schnepp, @FSCA, det PWM]
Lachnocrepis parallela TX
image
Laemostenus complanatus NV
(14)
Lebia abdominalis OK
(6)
Lebia arizonica [
verified? NV
image]
Lebia balli new to USA AZ
image
Lebia collaris OK
(6) SC[Insecta Mundi 0636:1-5, 2018]
Lebia fuscata OK
(6)
Lebia grandis ME
(10)
Lebia histrionica TX
image
Lebia lobulata KY
image
Lebia moesta CA [Placer Co., Roseville, 2008, leg/@GW Forister, det/PWM]
Lebia perita IA[Dallas Co: 2012 leg. E.Freese det/@FFP] NV
(14) TX[Jeff Davis Co: 2014 leg/det/@FFP]
Lebia pleuritica OK
(6) See challenge to OK record below by V.Belov 19 Sept 2018
Lebia rugatifrons (Chaudoir)
new to USA TX
image |
image
Lebia tricolor verified TX [Brazos Co & Sabine Co @TAMU det V.Belov v.i.]
Lebia variegata new to USA [under debate] AZ
image
Lebia 'UASM Lebia 53'
unresolved (= atypical color morph of
Lebia solea?) LA
image
Leptotrachelus dorsalis OK
(6) MA
image |
image WY
image
Leptotrachelus pallidulus MS["Hancock Co: det R.Davidson" per D.Hildebrandt pers. com. 02/20/13]
Lionepha australerasa sp. n.(18) CA
image
Lionepha chintimini →
L. erasa(18)
Lionepha "erasa" [Reconsider all of BugGuide's Lionepha in light of (18)]
Lionepha kavanaughi sp. n.(18)
Lionepha lindrothellus →
L. erasa (18)
Lionepha lindrothi sp. n.(18) CA
image
Lionepha lummi →
L. erasa(18)
Lionepha probata removed from synonymy with L. erasa(18) UT
image
Lionepha tuulukula sp. n.(18)
Lophoglossus vernix NC
image
Loricera pilicornis pilicornis NV
image(14)
Loxandrus → Oxycrepis (Loxandrus) by Will (2020)
(20); change epithets to feminine.
Medusapyga alsea sp. n. OR
image LaBonte & Madison 2023: p401
Medusapyga chehalis sp. n. WA
image LaBonte & Madison 2023: p401
Micratopus sp.
unresolved; M.?anaisae AZ
image
Micratopus aenescens WI[Rock Co., 2022, leg. L.Ferge, pinned M.Evans, det/@ PWM]
Micratopus insularis new to USA TX
image
Micratopus not-parviceps tentatively new to USA TX
image
Microlestes nigrinus verified NM [Santa Fe Co: 2007 leg. R. Delph, det/@ PWM]
Microlestes pusio VA
(9)
Mioptachys flavicauda DE
image
Mochtherus tetraspilotus SC [Insecta Mundi 0636:1-5, 2018]
Morion monilicornis KS
image
Myas coracinus ME
(10)
Nebria baumanni Kavanaugh
sp. n. NV[
Proc. Calif. Acad. Sci. 62:441-450 2015]
Nebria brevicollis BC [doi:10.4039/tce.2020.72] NY [v.i. CEJG comment/iNaturalist] TX[Smith Co/2015, Brazos Co/2015 & 2017, all @ TAMU, det V.Belov/D.Kavanaugh]
Nebria nivalis gaspesiana NH
(12)
Nebria suturalis UT
image
Nomius pygmaeus MO[Carter Co, Fremont, 1 ex @TAMU det V.Belov v.i.]
Notiobia cyanippa new to USA AZ
images
Notiobia nitidipennis KY
image
Notiobia terminata NV
(14) UT
image
Notiophilus aeneus AR
image
Notiophilus aquaticus NJ
image
Notiophilus biguttatus MA
images
Notiophilus nitens NV
(14)
Notiophilus semiopacus TX
image
Oodes americanus DC
(9) OK
(6)
Oodes brevis WI
(8)
Oodes fluvialis ND
image
Olisthopus brevicornis IN[Porter Co. det/@PWM]
Olisthopus iterans AR[Newton Co. det/@PWM]
Olisthopus micans ME
(10)
Omophron labiatum RI
image |
image
Omophron robustum MT
image
Omus c. californicus NV [Cicindela 53(2):25, 2021]
Ophonus puncticeps UT
image WA
image [The Coleopterists Bulletin 77(4): 606-608, 2023]
Oxycrepis (
Loxandrus)
cincinnati OK
(6)
Oxycrepis (
Loxandrus) nr.
circulus VA
(9)
Oxycrepis (
Loxandrus)
duryi OK
(6)
Oxycrepis (
Loxandrus)
erraticus OK
(6) VA[
Banisteria 45:61-62, 2015]
Oxycrepis (
Loxandrus)
nitidulus OK
(6)
Oxycrepis (
Loxandrus)
parallelus OK
(6)
Oxycrepis (
Loxandrus)
rectus OK
(6)
Oxycrepis (
Loxandrus)
robustus OK
(6)
Oxycrepis (
Loxandrus)
saphyrinus OK
(6)
Oxycrepis (
Loxandrus)
velocipes MI[Macomb Co, 1963, 1 ex @MSU, det PWM] OK
(6)
Panagaeus fasciatus ME
(10)
Paraclivina ferrea NE[Scotts Bluff Co: 2010 leg. FFP @J.Pretorius det FFP] VT
(12)
Paraclivina postica PA
image
Paraclivina striatopunctata MD[
Banisteria 38:71-84, 2011]
Paraclivina sulcipennis GA[Wheerler Co: 1♀ 2015 leg/det/@FFP]
Paradromius linearis genus & secies new to North America CA[Coleopterists Bull. 77(1):46, 2023] NS[Coleopterists Bull. 76(3):445, 2022]
Pasimachus sublaevis DE
image
Patrobus cinctus WI [Forest Co.,30 VI 2020, leg. L.Ferge, ♀ det/@PWM]
Patrobus fossifrons NV
(14)
Patrobus lecontei MT
image
Patrobus stygicus WY
under review image
Pentagonica bicolor AL
image GA[Wheerler Co: 1♀ 2015 leg/det/@FFP]
Pentagonica picticornis ME
(10)
Pentagonica maculicornis USA rec. n. FL [Monroe County, Big Pine Key, 16-17 V 2023, 2 exx. leg. D.Fine, det/@ D.Hildebrandt]
Perigona nigriceps MO
image
Philodes alternans KS
image
Philodes rectangulus NH
(12)
Philophuga viridis amoena SD
image
Phloeoxena signata TX[Angelina Co.: series @TAMU, det V.Belov v.i.]
Platynus acuminatus new to USA AZ[
image Cochise Co: 4 exx leg. C.Melton, det/@PWM] |
more images
Platynus opaculus VA
(9)
Platynus daviesi NH/VT
(12)
Platypatrobus lacustris MA
image
Plochionus timidus OK
(6) KY
image UT
image
Poecilus corvus AZ
image
Poecilus cyanicolor AZ
image
Poecilus lucublandus AZ
image
Polyderidius sp. AL, MS
see
Pogonodaptus mexicanus GA [Harden 2024: https://doi.org/10.1649/0010-065X-78.2.270]
Polyderis rufotestacea →
Nothoderis rufotestacea [genus n., Boyd & Erwin 2016] NV
(14)
Pseudamara arenaria → tribe Zabrini by
Gomez et al. 2016.
Pseudaptinus (Thalpius) dorsalis MO
image NC
image PA:
record by Messer 2011 overlooked by Bousquet 2012
Pseudaptinus (Pseudaptinus) lecontei NJ
image
Pseudaptinus (Thalpius) rufulus NV
(14) WA
(5)
Pseudaptinus (Thalpius) simplex new to USA AZ
images
Pseudomorpha huachinera sp. n. AZ
(4)
Pseudomorpha patagonia sp. n. AZ
(4)
Pseudomorpha penablanca sp. n. AZ
(4)
Pseudomorpha pima sp. n. AZ
(4)
Pseudomorpha santacruz sp. n. AZ
(4)
Pseudomorpha santarita sp. n. AZ
(4) NM
(4)
Pterostichus brevicornis NB
(24)
Pterostichus commutablis UT
image
Pterostichus corrusculus VT
(12)
Pterostichus escarinatus CO[Clear Crk. Co: 2014 leg/det/@FFP]
Pterostichus (Leptoferonia) hackerae sp. n. CA [K. Will 2016, Pan-Pacific Entomologist 92(4):177-188]
Pterostichus (Anilloferonia) lanei (Hatch, 1935) → replacement
Pterostichus (Anilloferonia) diana LaBonte, 2013 [Zootaxa 3682 (4):563-571]
Pterostichus luctuosus NV
(14)
Pterostichus permundus DC
(9) MD
image |
The Maryland Entomologist 7:35-47, 2017 VA
(9)
Pterostichus punctatissimus WI
(8)
Pterostichus restrictus UT
image
Pterostichus sculptus VA
(9)
Pterostichus subacutus AR[
Coleopt. Bull. 70(2):309-313, 2016]
Pterostichus tenuis VA
image(15)
Pterostichus vernalis VT
(12)
Rhadine caudata NC[first documented state record claimed by Riley & Browne in Coleopt. Bull. 68(2): 239-240, 2014. NC status given earlier by Bousquet (2012)]
Rhadine nivalis AZ
image |
image |
image
Scaphinotus elevatus flammeus MI["East Lansing", 1938, 1 ex @MSU, det W.Stehr & PWM]
Scaphinotus fissicollis WI
image
Scaphinotus hoodooensis sp. n. MT[
Proceed. Calif. Acad. Sci. 62:381-396, 2015]
Scaphinotus unistriatus SC
image
Scarites subterraneus NV
(14) UT
image
Scarites vicinus DC
(9) GA[Cherokee Co: det/@PWM] MD[
Banisteria 44:23-25, 2014] VA
(9)
Schizogenius depressus NV
(14)
Schizogenius lineolatus GA[Greene Co: 2013 leg/det/@FFP] MI[Macomb Co, 1964-1966, 8 exx @MSU, det W.Ruesink & PWM]
Selenophorus adjunctus →
Selenophorus granarius (22)
Selenophorus aequinoctialis sr. syn. of
S. famulus (22) AZ, CA, NV
Selenophorus balli sp. n. (22) TX
image
Selenophorus blanchardi FL
(22)
Selenophorus concinnus TX
(22)
Selenophorus contractus AL, FL
(22)
Selenophorus cupreolus →
Selenophorus pedicularius (22)
Selenophorus discoderoides →
Discoderus discoderoides (22) TX
images
Selenophorus discopunctatus AL
(22) GA
image
Selenophorus elongatus →
Selenophorus hylacis (22)
Selenophorus elytrostictus sp. n. (22) TX
image
Selenophorus fabricii sp. n. FL [
Zookeys 690: 1-195, 2017]
Selenophorus famulus →
Selenophorus aequinoctialis (22) NV
(14)
Selenophorus fatuus AZ
(22) KY
(22) image
Selenophorus gagatinus TN
(22)
Selenophorus granarius MI
(22), TN
(22), WI
(22)
Selenophorus houstoni →
Selenophorus pedicularius (22)
Selenophorus implicans →
Selenophorus pedicularius (22)
Selenophorus integer removed from FL (replaced in FL by
S. fabricii sp. n.)
Selenophorus intermedius new to USA TX
(22)
Selenophorus laesus →
Selenophorus palliatus (22)
Selenophorus maritimus AL
(22)
Selenophorus mexicanus new to USA TX
(22)
Selenophorus municeps →
Selenophorus ellipticus (22)
Selenophorus nanulus →
Selenophorus granarius (22)
Selenophorus neoruficollis sp. n. (22) LA, TX
Selenophorus nonellipticus sp. n. (22) AL, FL, GA, NC, SC
Selenophorus opalinus LA (see BugGuide & iNaturalist)
Selenophorus otiosus →
Selenophorus planipennis (22)
Selenophorus palliatus OK
(22) VA
verified image [also Sussex Co: 1♂ det PWM leg. R. Whalen @Virginia Polytech Institute; Virginia Beach: 1 ex. @VMNH det C.Harden & PWM]
Selenophorus palliatus OK, VA
(22)
Selenophorus pedicularius KY
(22) image MA
image QC
image
Selenophorus pumilus sp. n. (22) AZ
Selenophorus rileyi sp. n. (22) TX
Selenophorus riparius →
Selenophorus pedicularius (22)
Selenophorus scolopaceus →
Selenophorus aeneopiceus (22)
Selenophorus semirufus new to USA(22) AZ, CA, CO, NM, TX
Selenophorus seriatoporus new to USA(22) TX
image |
image |
image
Selenophorus subquadratus new to USA FL
(22)
Selenophorus subtropicus →
Selenophorus granarius (22)
Selenophorus undatus sp. n. (22) TX
Serranillus monadnock sp. n.(27) SC
image
Sericoda obsoleta OK
(6)
Somotrichus unifasciatus verified TX
image; VA[Powhatan Co: leg/det/@AV.Evans per C.Harden
(15)]
Sphaeroderus indianae AR
image
Stenocrepis duodecimstriata RI
image |
image |
image |
image
Stenocrepis mexicana MD[
The Maryland Entomologist 7:35-47, 2017]
Stenolophus comma NV
(14)
Stenolophus flavipes UT
image | UT [Garfield Co, May & Jun 2020, leg. D Uhey, det/@ PWM]
Stenolophus fuliginosus NV
(14) UT
image
Stenolophus humidus NB
(24)
Stenolophus lecontei ND
image
Stenolophus limbalis AZ[Mohave Co.: Pipe Spring NM 7/2012, 1♀ det/@PWM]
Stenolophus lineola ND
image WA
(5)
Stenolophus maculatus NM[Eddy Co: 2♀ 2014 leg/det/@FFP]
Stenolophus ochropezus ID?
image? ND
image NV
(14)
Stenolophus rugicollis AZ [Cochise Co., Miller Canyon, 2014, leg. CW Melton, det/@PWM]
Stenomorphus californicus rufipes verified for SC [v.i.
https://www.inaturalist.org/observations/62961506]
NC [Harden 2024: https://doi.org/10.1649/0010-065X-78.2.270]
Stolonis → Oxycrepis (Stolonis) by Will (2020)
(20); specific epithets stay feminine.
Stomis termitiformis CA
image
Synuchus impunctatus verified−CO[Animas Co: ♂♀ 2014 leg/det/@FFP] AR
image
Tachys (Paratachys) oblitus WI
(8)
Tachys (Paratachys) potomaca DC
(9)
Tachys (Paratachys) proximus MI[Counties Kalamazoo, Livingston, Midland, Van Buren, Wayne from 1890s - 1964 @MSU confirmed by PWM]
Tachys (Paratachys) scitulus RI
(13)
Tachys (Tachys) halophilus KS
image
Tachys (Tachys) mordax TX
image
Tachys (Tachys) vittiger TX(v.i.)
image
Tachyta angulata OK
(6)
Tachyta parvicornis ME
(10) OK[Latimer Co: 6 exx 1993-97 @TAMU det V.Belov v.i.]
Tetragonoderus intersectus OK
image
Tetragonoderus laevigatus LA
image image MS [Harrison Co.: Long Beach, 30.3397N/-88.1730W, 10 VI 2023, leg. R.Kergosien, det/@ DA Hildebrandt]
Tetragonoderus pallidus UT
image
Tetraleucus picticornis OK
image
Thalpius: see
Pseudaptinus
Trachypachus inermis NM
image, SD[as syn.
T. holmbergi in Coleopt. Bull. 66:357, 2012]
Trachypachus gibbsii UT
image
Trechus obtusus New to Eastern North America VA & NC [Coleop. Bull. 76(1): 61-69, 2022] WV
image
Trechus quadristriatus VA
image(15)
Trechus santaluciaensis sp. n. CA[Monterey Co., Donabauer 2014]
Trechus tenuiscapus NV
(14)
Trichotichnus fulgens RI
(13) image |
image |
image
Zuphiodes americanum CA
image
Zuphium → Zuphiodes for North America
(23).
Zuphium punctipenne Bates
new to USA TX[Hidalgo Co: det RL Davidson pers. com. 2012 @?TAMU] See
(23).
These claims for new geographic records require species confirmation in the opinion of PWM.
"
Poecilus cursitor" WA
image
"
Stolonis intercepta" OK
(6) and several other species listed 4/5/2014 in bold print are questionably assigned to "OK"
(6).
Items removed from the registry.
Bembidion cf. or nr. fortestriatum AZ [UW-IRC: Graham Co, det PWM;
Tree of Life website: Greenlee Co, det Maddison 2005] discussed below.
Trachypachidae & Rhysodidae: updated search for new records on BugGuide completed 7-Feb-2021.
A cited work that affects more than three species is assigned a number as follows.
1.
Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. Bousquet, Y. 2012. ZooKeys 245:1–1722.
2.
New and Additional Records of Ground Beetles and Wrinkled Bark Beetles for Virginia (Coleoptera: Carabidae, Rhysodidae). Roble, S.M. & R.L. Hoffman. 2012. Banisteria No 40:42-48.
3.
A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U.S.A. Hunting, W.M. 2013. ZooKeys 259: 1–73.
4.
Beetles that live with ants (Carabidae, Pseudomorphini, Pseudomorpha Kirby, 1825): A revision of the santarita species group. Erwin, T.L. & L.M. Amundson. 2013. ZooKeys 362:29-54.
5.
Ground beetles (Coleoptera, Carabidae) of the Hanford Nuclear Site in south-central Washington State. Looney, C., R.S. Zack, & J.R. LaBonte. 2014. Zookeys 396: 13-42.
6.
List of Coleoptera Collected in Latimer County, Oklahoma by Karl Stephan (2002). [Therein the uncertain species cited 4/5/2014 are marked in bold print]
7.
Life beneath the surface of the central Texan Balcones Escarpment: genus Anillinus Casey, 1918 (Coleoptera, Carabidae, Bembidiini): new species, a key to the Texas species, and notes about their way of life and evolution. Sokolov, I.M., J.R. Reddell, & D.H. Kavanaugh D.H. 2014. ZooKeys 417:71-101.
8.
Additions to the Checklist of Wisconsin Ground Beetles (Coleoptera: Carabidae) including Adventive Harpalus rubripes (Duftschmid) among Seven New State Records. Messer, P.W. 2014. The Great Lakes Entomologist 47: 66-72.
9.
Twelve Ground Beetles New to Virginia or the District of Columbia and an Annotated Checklist of the Geadephaga (Coleoptera, Adephaga) from the George Washington Memorial Parkway. Steury, B., & P.W. Messer. 2014. Banisteria 43:40-55.
10.
The Ground Beetle (Coleoptera: Carabidae) Fauna of Maine, USA. Dearborn, R.G., R.E. Nelson, C. Donahue, R.T. Bell, & R.P. Webster. 2014. The Coleopterists Bulletin 68: 441 - 599.
11.
Five new species of Anillinus Casey from Alabama with a key to the Alabama species (Carabidae: Trechinae: Bembidiini). Sokolov, I.M. 2012. Annals of Carnegie Museum 81:61-71.
12.
Carabidae of Vermont and New Hampshire. Bell, R.T. 2015. Shire Press.
13.
The Beetle Fauna of Rhode Island. Sikes, D.S. 2004. [NB: Some of the RI records in Sikes' well-documented 2004 list were likely omitted by Bousquet (2012) because they were based on very old specimens identified before modern species keys became available.]
14.
Additions to the knowledge of Nevada carabid beetles (Coleoptera: Carabidae) and a preliminary list of carabids from the Great Basin National Park. Will, K., Madan, R., Hsu, H. 2017. Biodiversity Data Journal 5: 1- 23.
15.
Eight Ground Beetles (Coleoptera: Carabidae) New to Virginia, with Additional Records for West Virginia and Maryland. Harden, C.W. 2018. Banisteria 50: 15 - 20.
16.
Range update for eight species of the subfamily Elaphrinae (Coleoptera: Carabidae) in Ontario's Far North & Nunavut, Canada. Fleming, K.J. & D.V. Beresford. 2019. The Coleopterists Bulletin 73(2): 433 - 439.
17.
Unifying systematics and taxonomy: Nomenclatural changes to Nearctic tiger beetles (Coleoptera: Carabidae: Cicindelinae) based on phylogenetics, morphology and life history. Duran, DP, & HM Gough. 2019. Insecta Mundi 0727: 1 - 12.
18.
Species delimitation, classical taxonomy and genome skimming: a review of the ground beetle genus Lionepha (Coleoptera: Carabidae). Maddison, D.R. & JS Sproul. 2020. Zoological Journal of the Linnean Society XX: 1 - 46.
19.
Four new species of the genus Anillinus Casey (Coleoptera, Carabidae, Anillini) from Alabama, U.S.A., with a revised key to the Alabama species. Sokolov, I.M. 2020. Zootaxa 4808(3): 547-559.
20.
Phylogeny and classification of the genus-group taxa of Loxandrina (Coleoptera, Carabidae, Abacetini). Will, K. 2020. Dtsch Entomol Z 67: 151-182.
21.
Species delimitation in the ground beetle subgenus Liocosmius (Coleoptera: Carabidae: Bembidion), including standard and next-generation sequencing of museum specimens. Maddison, D.R. & K.W. Cooper. 2014. Zoological Journal of the Linnean Society, 172: 741–770.
22.
A review of Nearctic Selenophorus Dejean (Coleoptera: Carabidae: Harpalini) north of Mexico with new species, new synonyms, range extensions, and a key. Messer, P.W. & B.T. Raber. 2021. The Coleopterists Bulletin, 75(1): 9-55. URL:
https://doi.org/10.1649/0010-065X-75.1.9
23.
Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi, new species, and Zuphioides, new genus, and classification of the genera (Coleoptera, Carabidae). Ball, G.E. & D. Shpeley. 2013. ZooKeys 315: 17-54.
24.
New Coleoptera records from eastern Canada, with additions to the fauna of Manitoba, British Columbia, and Yukon Territory. Webster, RP., P. de Tonnancour, JD. Sweeney, VL. Webster, CA. Kostanowicz, C. Hughes, RS. Anderson, J. Klymko, C. Chantal, R. Vigneault. 2020. ZooKeys 946: 53-112.
25.
Classification of the genus Harpalus (Coleoptera, Carabidae) of the World based on imaginal morphology. Kataev, B.M. 2023. Diversity 15, 971. [BugGuide's classification of
Harpalus will be adjusted according to this revision.]
26.
Revision of the elongatus and pecki species groups of Anillinus Casey, 1918 (Coleoptera: Carabidae: Trechinae: Anillini), with four new species and notes on Anillinus turneri Jeannel. Harden, C.W. & M.S. Caterino. 2024. Insecta Mundi 1057: 1-31.
27.
Systematics and biogeography of Appalachian Anillini, and a taxonomic review of the species of South Carolina (Coleoptera, Carabidae, Trechinae, Anillini). Harden, C.W. & M.S. Caterino. 2024. ZooKeys 1209: 69-197.
