Found at lights on 2nd night of the 2013 BugGuide summer gathering.
When I photographed this fly I figured it was the same as the other pyrgotid flies we were seeing that night, some of which are posted below:
However at full-size on the computer screen this one looked different...with its the strongly banded tibiae, and more "blocky" mouthparts with black palps. I initially guessed those differences might be due to sexual dimorphism within the same species. But they turned out to be different species. Working through the keys in the MND
(1) and
Steyskal (1978) this individual went to genus
Boreothrinax, with a crucial character being the distinctively constricted basal 1/3 of the hind tibia.
The genus
Boreothrinax was erected in Steyskal (1978) with three species: two were new species,
B. shewelli and
B. dichaetus; and the third,
B. maculipennis, was moved from
Sphecomyiella maculipennis (originally
Oxycephala maculipennis Macquart 1846). Using the key to species in Steyskal (1978), I arrived tentatively at
B. maculipennis, based on the combination of Arizona location and bare prosternum (
B. dichaetus is also recorded from AZ, but has prosternum with fine lateral setae). Unfortunately the description of
B. maculipennis in Steyskal is minimal so I decided to search for more info on the web.
When I checked BugGuide I found a number of images, but was surprised to see them posted under the name
"Pyrgota" maculipennis, with the
info page listing
Boreothrinax maculipennis as a synonym. The initial BugGuide post under
P. maculipennis was made in 2005 and is shown in the thumbnail below:
In that post Gary Steck (again, in 2005) made the ID of
Pyrgota maculipennis, noting the shape and banded color pattern of the hind tibia, and mentioning that
B. maculipennis was a synonym. A species page was made and all subsequent posts have been placed under
Pyrgota maculipennis.
But why did the genus name
Pyrgota get substituted for
Boreothrinax here? Was there a publication sometime after the 1987 MND that had presented a basis for making
B. maculipennis a junior synonym of
Pyrogta maculipennis? If so, I figured that would be reflected in the more recent Manual of Central American Diptera
(2) Vol 2, published in 2010. But the MCAD recognizes the same 3 US species of
Boreothrinax as the MND
(1) did...namely those circumscribed in Steyskal (1978), which is cited by both the MND and the MCAD as their reference for that genus (as well as the US species of
Pyrgota). Other published references I searched out, like the 1996 paper
here also used the name
Boreothrinax maculipennis.
While I was unable to find any actual reference papers addressing whether
Boreothrinax maculipennis was transferred to
Pyrgota and made a junior synonym, there
were numerous Google hits for the name
Pyrgota maculipennis. These were principally in the form of the "web robot" type taxonomic names web sites which have proliferated in recent years (e.g.
Diptera Nomenclator,
ITIS,
GNI,
EOL,
ADW,
Gwannon, etc.), as well as Flickr posts and other photo posts which appeared to get the name
Pyrgota maculipennis from BugGuide...perhaps in combination with the robotic taxonomy sites.
Most of the robotic taxonomy sites offer little content other than names and often seem to simply copy information from each other and lack logical consistency. For instance, the ITIS web site lists
Pyrgota as a genus having 6 valid species...including the 3 species
dichaetus, maculipennis, and
shewelli that Steyskal put in
Boreothrinax. They also include
valida among those 6
Pyrgota species names, apparently referring to
Sphecomyiella valida, which they also list as a valid species on their
Sphecomyiella web page...so they are apparently asserting two different "valid" names for the same species! I also noticed that the
ITIS Boreothrinax web page indicates
Boreothrinax is a valid genus...but they show it as having no species!! Meanwhile, there is no mention on the
ITIS page for Pyrgota maculipennis of the purported synonym
Boreothrinax maculipennis.
Compare the situation for the ITIS listings discussed above to the
BugGuide Pyrgota info page, created in 2004 (i.e. before the
Pyrgota maculipennis guide page was created in 2005), which cites the now defunct
Nearactica.com web site as listing just 2 species in the genus:
fenestrata and
undata. That agrees with the treatment in Steyskal (1978), which appears to still be valid as far as I can tell.
All this makes the names given on the "web robot" pages seem fishy...which it is consistent with the reservations expressed by various taxonomists I know regarding the accuracy of some (but not all) such listings.
So...is anyone aware of a bonafide post-1987 publication or catalog indicating that
Boreothrinax maculipennis was made a junior synonym for
Pyrgota maculipennis?
My guess is that, unless I'm missing something, it appears that
Boreothrinax is the valid genus name here, in which case the genus name currently being used for this species on BugGuide should be corrected.