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Genus Anatis - Giant Lady Beetles

Eye-spotted Lady Beetle - Anatis mali Anatis (I think) - Anatis Eye-spotted Lady Beetle - Anatis mali Anatis labiculata (Say) - Anatis labiculata Anatis rathvoni, I think - Anatis rathvoni Anatis labiculata? - Anatis labiculata Eyed-spotted Lady Beetle - Anatis mali - Anatis mali ID help please - Anatis lecontei
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga
No Taxon (Series Cucujiformia)
Superfamily Coccinelloidea
No Taxon (Coccinellid group)
Family Coccinellidae (Lady Beetles)
Subfamily Coccinellinae
Genus Anatis (Giant Lady Beetles)
Synonyms and other taxonomic changes
Myzia LeConte 1852 (in part)
Explanation of Names
Anatis Mulsant 1846
Numbers
4 spp. in our area, 6 total
Size
7.2-10.5 mm(1); the largest coccinellids of our fauna
Identification
see (1)
Patch of fine, short hairs at rear of each elytron. Antennae with 10 segments, terminating in triangular club. Meso- and metatibia each with 2 spurs. Tarsal claw with large, subquadrate basal tooth.
Range
in NA, across the north (AK-NF)(2) to NM-SC
Habitat
Usually arboreal
Food
Mostly aphids, but also caterpillars and sawfly larvae, especially those in trees.
Remarks
Adults darken with age. The oldest individuals may appear to have no black spots because the base color is so dark.
Print References
McKenzie H.L. (1936) An anatomical and systematic study of the genus Anatis of America. U. Cal. Publ. Entomol. 6(10): 263-272.
Works Cited
1.The Coccinellidae (Coleoptera) of America North of Mexico
Robert D. Gordon. 1985. Journal of the New York Entomological Society, Vol. 93, No. 1.
2.Checklist of beetles (Coleoptera) of Canada and Alaska. Second edition
Bousquet Y., Bouchard P., Davies A.E., Sikes D.S. 2013. ZooKeys 360: 1–402.