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Genus Anatis - Giant Lady Beetles

Eye-spotted lady beetle - Anatis mali Flying saucer ladybug - Anatis rathvoni Anatis (I think) - Anatis Possible Eye-spotted Lady Beetle pupa - Anatis large ladybird larvae - Anatis Fifteen-spotted Lady Beetle - Anatis labiculata fifteen-spotted lady beetle pupa - Anatis labiculata Lady Beetle - Anatis
Classification
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
No Taxon (Series Cucujiformia)
Superfamily Coccinelloidea
No Taxon (Coccinellid group)
Family Coccinellidae (Lady Beetles)
Subfamily Coccinellinae
Genus Anatis (Giant Lady Beetles)
Synonyms and other taxonomic changes
Myzia LeConte, 1852 (in part)
First described in 1846 by Étienne Mulsant
Numbers
6 species worldwide, with 4 species in North America:
Anatis labiculata Say 1824 (Coccinella)
Anatis lecontei Casey 1899 (Anatis)
Anatis mali Say 1825 (Coccinella)
Anatis rathvoni LeConte 1852 (Myzia)
Size
Length 7.2 to 10.5 mm, width 5.5 to 9.0 mm.(1)
Identification
Gray, yellowish, red, or purple (darkening with age).
Pronotum (between wings and head) black with prominent white markings.
Oval or shield-shaped, sometimes with angular rather than rounded outline.
Strongly to weakly explanate (helmet-shaped) in profile.
Patch of fine, short hairs at rear of each elytron (wing cover).
Antennae with 10 segments, terminating in triangular club.
Apex of middle and hind tibia each with 2 spurs.
Tarsal claw with large, subquadrate basal tooth.
Range
Ontario to South Carolina, west to southern Alaska and New Mexico.
Habitat
Usually arboreal (in trees) in forests and woods.
Food
Mostly aphids, but also caterpillars and sawfly larvae, especially those in trees.
Remarks
This genus contains the largest lady beetles in North America.
All species darken with age. The oldest individuals may appear to have no black spots because the base color is so dark.
Print References
Gordon, 1985, especially pp. 752-764, figures 614-625.(1)
McKenzie, H.L. 1936. An anatomical and systematic study of the genus Anatis of America. University of California Publications in Entomology 6(10): 263-272.
Internet References
Histoire naturelle des Coleopteres de France: Sulcicolles-Securipalpes (1846), p.133 - Mulsant's original description of the genus (in French)
Works Cited
1.The Coccinellidae (Coleoptera) of America North of Mexico
Robert D. Gordon. 1985. Journal of the New York Entomological Society, Vol. 93, No. 1.