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Genus Tomarus


Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae) new classification and taxonomic revision
By López-García, M.M. and C. Deloya
Zootaxa 5211(1): 1-119., 2022
Abstract and References - Zootaxa

López-García, M.M. and C. Deloya. (2022) Cladistic analysis reveals polyphyly of Tomarus (Coleoptera: Scarabaeidae: Dynastinae): new classification and taxonomic revision. Zootaxa 5211(1): 1-119.

Abstract

The phylogeny of Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) was explored by parsimony using morphological characters to approach its taxonomic and nomenclature problems. The genus was not supported as a monophyletic group, and the recovered lineages did not match at all any of the previously known classifications. Three different lineages and six monophyletic groups were recovered, which could be treated as independent genera. However, the classification proposed here is as conservative as possible concerning to the creation of many new names and generic taxa.

Scarab beetles of the genus Bothynus in the United States (Coleoptera: Scarabaeidae)
By O. Cartwright
Proceedings of the United States National Museum, 108(3409): 515-541, 1959
Online here.

Monograph of the genus Tomarus (=Bothynus, Ligyrus)

Collagenus dasysternus ... with a key to the New World genera of Pentodontini
By Brett Ratcliffe and Martin Hardy
The Coleopterists Bulletin, Vol. 59, pp.143-150, 2005
Brett Ratcliffe and Martin Hardy, Collagenus dasysternus, a new genus and species of Dynastinae from eastern Venezuela with a key to the New World genera of Pentodontini (Coleoptera: Scarabaeidae: Dynastinae), The Coleopterists Bulletin 59(1): 143-150, 2005.

From BG's Pentodontini Info tab, "One error noticed; couplet 11- range of genus Aphonus should be 'United States east of the Rocky Mountains'".

Full text

Taxonomic review of the North American dung beetle genus Melanocanthon Halffter, 1958
By Edmonds, W.D.
Insecta Mundi, 2023
Edmonds, W.D. 2023. Taxonomic review of the North American dung beetle genus Melanocanthon Halffter, 1958 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini). Insecta Mundi 1014: 1-28.

New country and state Records of Scarabaeidae (Coleoptera) in the USA.
By Warner, W.B.
The Coleopterists Bulletin 76(1): 128-129., 2022
BioOne

Warner, W.B. (2022). New country and state Records of Scarabaeidae (Coleoptera) in the USA. The Coleopterists Bulletin 76(1): 128-129.

This brief paper describes new Arizona state distribution records for Haroldiellus sallei (Harold) (Scarabaeidae: Aphodiinae: Aphodiini) and Phyllophaga crinita (Burmeister) (Scarabaeidae: Melolonthinae: Rhizotrogini), as well as new USA country records (Arizona and Texas, respectively) for Haroldiataenius limbatus (Bates) (Scarabaeidae: Aphodiinae: Eupariini) and Amithao erythropus (Burmeister) (Scarabaeidae: Cetoniinae: Gymnetini). Vouchers of all the species are deposited in the Hasbrouck Insect Collection at Arizona State University, Tempe, Arizona, USA.

The species of Scarabaeidae (Coleoptera) associated with sugarcane in south Florida
By R.D. Gordon and D.M. Anderson
The Florida Entomologist, 64 (1): 119-138 , 1981
"Those species of Scarabaeidae known to be associated with sugarcane in the Lake Okeechobee area of Florida are discussed and delineated. The species are: Phyllophaga latifrons (LeConte), Anomala marginata (F.), Cyclocephala parallela Casey, Dyscinetus morator (F.), Ligyrus subtropicus (Blatchley), and Euphoria sepulchralis (F.). Descriptions and illustrations of the larvae of all these species are presented as well as descriptions and illustrations of the adults of all species except Euphoria sepulchralis (F.). A key to the south Florida species of the genus Cyclocephala is also included."

A catalog of the Coleoptera of America North of Mexico: Family Scarabaeidae, Subfamilies Rutelinae and Dynastinae
By Hardy A.
USDA Agric. Handbook 529-34b, 57 pp., 1991

The Scarab Beetles of Florida
By Robert Woodruff
Florida Department of Agriculture and Consumer Services, 1973
Full title: Arthropods of Florida and Neighboring Land Areas. Volume 8. The Scarab Beetles of Florida (Coleoptera: Scarabaeidae). Part I. The Laparosticti (Subfamilies: Scarabaeinae, Aphodiinae, Hybosorinae, Ochodaeinae, Geotrupinae, Acanthocerinae). Part II covers May Beetles (Phyllophaga).

Has black-and-white photos. Apparently out of print, but still available from some suppliers--see comments. Also available on-line as part of:
The Arthropods of Florida--scroll down to Vol. 8 and 13 - Scarab Beetles of Florida