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For Insects, Spiders & Their Kin
For the United States & Canada

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Family Platypezidae - Flat-footed Flies

Representative Images

Flat-footed Fly - Polyporivora Platypezidae? - Lindneromyia flavicornis Platypezid - Platypeza anthrax - female flat-footed fly! dorsal Lindneromyia flavicornis - female Lindneromyia flavicornis - female Calotarsa Fly 3-5mm - Callomyia

Classification

Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Diptera (Flies)
No Taxon ("Aschiza")
Superfamily Platypezoidea
Family Platypezidae (Flat-footed Flies)

Other Common Names

Smoke Flies - because they are fond of wood smoke(1)

Synonyms and other taxonomic changes

Genera treated in the Opetiinae in (2) now belong to 2 diff. subfamilies, Melanderomyiinae and Microsaniinae; the Palaearctic genus Opetia is now considered to be a sole member of its own family, Opetiidae

Explanation of Names

Platypezidae Latreille 1829

Numbers

4 subfamilies, with ~70 spp. in 17 genera in our area(3) and ~300 spp. in ~30 genera total(4)

Size

Mostly 2‒5 mm (Calotarsa 6‒10 mm)

Range

Mostly n. hemisphere, incl. much of NA

Season

Late summer‒fall

Food

Adults may feed on fungi. Some groups take honeydew from leaves(5)

Remarks

Males sometimes swarm, and females are attracted to these swarms. Some are attracted to smoke. Larvae live on fungi, some genera are associated with stinkhorn mushrooms. The elongated tarsi may be an adaptation for spreading host spores. Have a strong propensity to form swarms. Maybe the smell of smoke triggers them to anticipate fire-loving fungi growth that occur on charred remains after a forest fire.(1)

Print References

(6)(7)

Works Cited

1.Hidden Company that Trees Keep: Life from Treetops to Root Tips
James B. Nardi. 2023. Princeton University Press.
2.Manual of Nearctic Diptera Volume 2
Varies for each chapter; edited by J.F. McAlpine, B.V. Petersen, G.E. Shewell, H.J. Teskey, J.R. Vockeroth, D.M. Wood. 1987. Research Branch Agriculture Canada.
3.American Insects: A Handbook of the Insects of America North of Mexico
Ross H. Arnett. 2000. CRC Press.
4.Catalogue of Life
5.The flat-footed flies: (Diptera: Opetiidae and Platypezidae) of Europe
Chandler P.J. 2001. Fauna Entomologica Scandinavica 36. Brill Academic Pub. 278 pp.
6.A review of the Platypezidae of Eastern North America
Johnson, C.W. 1923. Occasional Papers of the Boston Society of Natural History, 5: 51-58 + plate.
7.A revision of the genera of Platypezidae, with the descriptions of five new genera, and considerations of phylogeny...
Kessel E.L., Maggioncalda E.A. 1968. Wasmann J. Biol. 26: 33-106.