There is fairly strong molecular evidence, for instance as indicated by specimens examined through
BOLD Systems, that two genetically distinct groupings are involved here. However, at this stage it seems unclear how to differentiate them short of nucleotide analysis, and there is need of further analysis and correlation of information. Also, their relationshipe to other species (notably to
P. ursaria) needs clarified. It is pretty much certain that both groups are represented by the specimens posted here. As things stand now, it seems impossible to separate the two groups reliably based on color pattern alone, and for now they are grouped together here under the one name. Those specimens previously stated to be a match for
P. californiaria here and also at BOLD, are part of one of these two groups, and do not appear to actually belong with that name. Hopefully better understanding along with clarification will come soon.