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Family Ochodaeidae - Sand-loving Scarab Beetles

Neochodaeus frontalis Parochodaeus - Parochodaeus inarmatus Neochodaeus Sand-lover Ochodaeidae Ochodaeidae, Parochodaeus sp. - Parochodaeus Beetle, anterior-lateral, showing frontal tubercle on the head behind the anterior margin - Neochodaeus frontalis Madera Canyon-0BG.J.2016.02 - Xenochodaeus
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Coleoptera (Beetles)
Suborder Polyphaga (Water, Rove, Scarab, Long-horned, Leaf and Snout Beetles)
Superfamily Scarabaeoidea (Scarab, Stag and Bess Beetles)
Family Ochodaeidae (Sand-loving Scarab Beetles)
Explanation of Names
Ochodaeidae Mulsant & Rey 1871
ca. 80 spp. in 12 genera worldwide(1), 22 spp. in 6 genera in NA as follows(2):
Family OCHODAEIDAE Mulsant & Rey 1871
Subfamily OCHODAEINAE Mulsant & Rey 1871
Codocera Eschscholtz 1821:398
gnatho (Fall in Fall & Cockerell 1907)
Cucochodaeus Paulsen 2007
sparsus (LeConte 1868:51)
Neochodaeus Nikolajev 1995
frontalis (LeConte 1863)
praesidii (Bates 1887)
repandus (Fall 1909)
striatus (LeConte 1854)
Parochodaeus Nikolajev 1995
biarmatus (LeConte 1868)
californicus (Horn 1895)
duplex (LeConte 1868)
*howdeni (Carlson 1975)
inarmatus (Schaeffer 1906)
pectoralis (LeConte 1868)
*peninsularis (Horn 1895)
pixius Paulsen 2011
*ritcheri (Carlson 1975)
Xenochodaeus Paulsen 2007
americanus (Westwood 1852)
*luscinus (Howden 1968)
musculus (Say 1835)
*planifrons (Schaeffer 1906)
*simplex (LeConte 1854)
*ulkei (Horn 1876)
Subfamily CHAETOCANTHINAE Scholtz 1988
Pseudochodaeus Carlson & Ritcher 1974
estriatus (Schaeffer 1906)
the beetles have prominent mandibles visible from above; mesotibia with long spur serrate along one edge:
worldwide except Australia, in temperate and tropical habitats; in our area, most diverse in the southwest
One southeastern species is in sandy, wooded habitats. Another eastern species makes burrows in hard packed clay in wooded areas and can be found on hiking trails. Western species generally in sandy habitats. Species are generally attracted to light.
Unknown. Larvae of most species have never been found.
See Also
female Odonteus are superficially similar but have bare dorsum and divided eyes
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