Identification, Images, & Information
For Insects, Spiders & Their Kin
For the United States & Canada
Clickable Guide
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Family Aradidae - Flat Bugs

A flat bug - Aradus robustus Which Flat Bug? - Aradus quadrilineatus Pine Flat Bug - Aradus kormilevi - male Mezira - Mezira subsetosa Aradidae - Aradus Neuroctenus - Neuroctenus elongatus - female flat bug - Aradus crenatus Small bug, possibly a Flat Bug (Aradidae )
Kingdom Animalia (Animals)
Phylum Arthropoda (Arthropods)
Subphylum Hexapoda (Hexapods)
Class Insecta (Insects)
Order Hemiptera (True Bugs, Cicadas, Hoppers, Aphids and Allies)
Suborder Heteroptera (True Bugs)
Infraorder Pentatomomorpha
Family Aradidae (Flat Bugs)
Explanation of Names
Aradidae Spinola 1837
~130 spp. (>⅔ of which are in Aradus) in 13 genera in our area, arranged in 5 subfamilies(1)(2); close to 2,000 spp. in 230 genera of 8 subfamilies worldwide(3)
Overview of our fauna (* –taxa not yet in the guide)
Subfamily Aradinae Aradus · Quilnus
Subfamily Calisiinae Calisius
3‒20 mm (our spp. ≤11 mm)
Key to genera and the commonest eastern spp. in (5) • to northern spp., in (6) • to Michigan spp., in (7)
Many nw. species imaged at (8)
somewhat helpful key to genera spread via human agency in (9)
worldwide; most diverse in Australia(3)
mostly under bark, on fallen logs & tree limbs, bracket fungi; some smaller forms in leaf litter
Mycelia or fruiting bodies of wood-rotting fungi(3)
Life Cycle
many bark-dwelling spp. are gregarious(3)
mouthparts at rest coiled inside the head
Print References
Works Cited
1.State records, confirmations, and habitats of Aradidae (Hemiptera: Heteroptera) from Louisiana, U.S.A.
Taylor S.J., Gil S.A. 2009. Fla. Entomol. 92: 199-207.
2.Taylor S. (2002-2011) Aradidae (Flatbugs)
3.Zoological catalogue of Australia: Hemiptera: Heteroptera (Pentatomomorpha)
Cassis G., Gross G.F. 2002. CSIRO Publishing, 751 pp.
4.New Florida and United States Heteropteran records (Hemiptera: Aradidae, Coreidae, Miridae)
Henry T.J., White C.E., Halbert S.E. 2021. Proc. Ent. Soc. Wash. 123: 538-550.
5.How to Know the True Bugs
Slater, James A., and Baranowski, Richard M. 1978. Wm. C. Brown Company.
6.The Aradidae of Canada (Hemiptera: Aradidae)
Matsuda R. 1977. The insects and arachnids of Canada, Pt. 3. Ottawa: Agriculture Canada. 116 pp.
7.A synopsis of the Flat Bugs (Heteroptera: Aradidae) of Michigan
Swanson D.R. 2020. Great Lakes Entomol. 53: 83‒104.
8.Photo gallery of the Spencer Entomological Collection
9.Flat Bugs (Hemiptera: Heteroptera: Aradidae) intercepted at United States ports of entry
Smith-Pardo A., Beucke K. 2015. Pan-Pac. Entomol. 91: 58‒81.
10.Aradidae (Heteroptera) new to North America and Canada, with some additional provincial and state records for Canada and the USA
Heiss E., Scudder G.G.E. 2019. Linzer biol. Beitr. 51: 821‒829.
11.New state records for some flat bugs (Heteroptera: Aradidae) of the United States
Swanson D.R. 2020. Great Lakes Entomol. 53: 105‒108.
12.Annotated checklist of the Aradidae (Hemiptera: Heteroptera) or "Flat Bugs" of Iowa
Freese E., Chordas S.W. 2022. Great Lakes Entomol. 55: 110‒115.
13.Aradidae (Hemiptera: Heteroptera) emergent from coarse and fine woody debris in Louisiana
Ferro M.L., Swanson D.R., Ulyshen M.D. 2023.